Poa iconia var. pelasgis Soreng, Castanea

1.2.Poa iconia var. pelasgis(H. Scholz) Soreng, Castanea 83 (2): 271. 2018. Fig. 10 A – C = Poa eigii Feinbrun, Bull. Misc. Inform. Kew 1940: 280. 1941. Type. Palestine, Judean Desert: km. 18 on the Jerusalem-Jericho road, grey compact and gravelly soil, N-NW exp., 28 Mar 1935, A. Eig, M. Zohary et A. Grizi s. n. (lectotype, here designated: HUJ- 29079 [HUJ 25031501. jpg. image!]. Note. Original material at K [K 000641323] annotated as “ cotype! ”, and later as isotype, is a syntype collected on 22 Mar 1835). = Poa sinaica subsp. graeca H. Scholz, Bot. Chron. (Patras) 3 (1–2): 20. 1983 b. Type specimen. Peloponnes, Nom. Lakonis, between the villages of Chania and Krokee, c. 24 km SSE Sparti, along main road to Nonemvasia, 250 m, olive grove and road embankment, 29 Mar 1980, Strid et Baden 71334 a (holotype: C {n. v.}). ≡ Poa pelasgis H. Scholz, Willdenowia 15 (1): 94 (1985). Type. Karpathos • Kalilimni, SE-Flanke, 35 ° 35 ' 05 " N, 27 ° 08 ' E, Felsbänder, Spalten und feinerdegefüllte Dolinen in Kleinstrauchgarrigues über verkarstetem Kalkgestein, 700–950 m, 8 May 1982, Greuter, Greuter, Pleger et Raus WG- 18935, TR 6247. (holotype: B [B 100508138]). Description. Perennials; hermaphroditic; tufted; primary roots 0.1–0.2 mm diam., tufts without rhizomes or stolons, dense, 1–9, av. 5 cm tall, forming mats of bulbs, light to dark green, sometimes violaceous in part; tillers intravaginal, bulbous at the base, erect; intravaginal prophylls 0.5–2 cm long, mostly included in the adjacent sheaths, keels smooth or distally antrorsely scabrous to scaberulous; bulbs 0.4–1.5 cm long, 1.3–2.3 mm wide, subglobose to pyriform, usually erect, slender, the outer sheaths papery to subcoriaceous, weakly veined, slender veined, persisting as a tunic, inner basal sheaths thickened, indurate at the base for 2–5 mm. Culms (10 –) 15–45, av. 28 cm tall, slender, erect or geniculately ascending, internodes smooth or scabrous, glabrous or hispidulous to strigulose below the lower nodes, 1 (– 2) nodes exerted, uppermost node at ca 1 / 3 plant height. Leaves mostly basal, culm leaves 1–2; leaf sheaths strongly imbricated at base, keel scabrous sometimes scaberulous on the sides near the collar, or sparsely to densely hispidulous to strigulose (hairs to 0.15 mm long); flag-leaf sheaths (2.4 –) 4.5–10 (– 12), av. 7.5 cm long, margins fused 10–20 % the length, smooth or lightly scaberulous, glabrous; throats and collars of some basal or lower culm leaves commonly scaberulous or hispidulous to strigulose on the sides or only near the margins; ligules of upper culm leaves 1–3 (– 4) mm long, scarious, watery-white, abaxially smooth, scabrous, or strigulose. apices obtuse to acute; ligules of sterile shoots 0.2–1 mm long, not or narrowly decurrent (without inflated cells at collar margins), apices truncate to acute, often dentate, abaxially scabrous; blades of culm 2–5 (– 6) cm long, 0.2–0.5 mm wide as folded, 0.4–1 mm wide; flag-leaf blades (0.4 –) 1.3–4 (– 6), av. 2.5 cm long, av. 1 / 3 the length of its sheath, blades equal or gradually reduced upward, folded, slender to filiform, thin, flaccid or retaining shape in age, abaxially proximally scabrous along the keel and sometimes the surfaces, margins scabrid, or keel, surfaces, and margins hispidulous-strigulose, apices narrowly prow-tipped; sterile shoot blades 1–10, av. 5 cm long, filiform, withering in age. Panicles (1.5 –) 3–7 cm long, with (20 –) 30–70 spikelets, erect, narrowly ovoid to lanceolate, loosely contracted (open in anthesis), crowded or loose; peduncles 7–15 cm long, smooth, proximal internodes ca 0.8–1 cm long; branches (2 –) 3–4 (– 5) at the lowest node; primary branches (0.5 –) 1–3, with (3 –) 6–12 spikelets in the distal 1 / 3–1 / 2, steeply or laxly ascending, terete to weakly angled, slender to subcapillary, sparsely to moderately scabrous; lateral pedicels mostly <1 mm long. Spikelets (2.7 –) 3–4 mm long, narrowly ovate, compact, laterally compressed, light to dark green, often violaceous in part; not bulbiferous, spikelets all normal; florets 2–3 (– 5); rachilla internodes short, mostly 0.4–0.7 mm, obscurely muriculate, glabrous; glumes subequal, lanceolate, sub-lustrous, distinctly keeled, distally moderately scabrous, edges sparsely to moderately scaberulous; apices acute to narrowly acute; lower glumes 1.8–2.8 mm long, usually 1.2–2 x narrower than the upper glume, 1 - veined, narrowly lanceolate; upper glumes (1.8 –) 2.3–3 mm long, lanceolate; callus glabrous; lemmas (2 –) 2.2–3.2 mm, 5 - veined, lanceolate to broadly lanceolate, light green, commonly violaceous, sub-lustrous, distinctly keeled, keels and marginal veins pectinate-ciliate, longer hairs 0.5–1 mm long, single file, not crowded (sharply pointed), between veins smooth, glabrous, intermediate veins faint to obscure, margins inrolled below, distally scarious / hyaline, edges moderately scaberulous, apices acute to narrowly acute, pointed; palea keels scabrous in distal 2 / 3–1 / 2, between keels smooth, glabrous. Flowers bisexual; lodicules 0.5 mm long, broad, lobed; anthers (1 –) 1.2–1.5 mm long. Caryopsis 1.3–1.4 mm long, trigonous, broadly sulcate, hard, adherent to the palea; hilum ca 0.2 mm long, elliptical. 2 n = polyploid, of uncertain application. Clade code: Nn. Distribution and ecology. Croatia; Greece (including Type of Poa pelasgis); Cyprus; Israel (including Type of Poa eigii); Jordan; Turkey. Mediterranean upland forest openings to subalpine; in Turkey, 200–1,600 m. Conservation status. None. Comments. Scholz (1985) equated Poa sinaica subsp. graeca with P. pelasgis when describing the latter with a new Type. Poa eigii is indistinct from var. pelasgis, except perhaps for the greater tendency to form large mats. Feinbrun (1986 a) noted a rare glabrous form and a bulbiferous form from Jordan but did not effectively name those. One specimen from the Negev Desert det. as P. eigii by Scholz, also exhibits the loosely strigulose hairs along its lower sheaths, characteristic of many P. iconia specimens. Scholz (1985) did not provide any distinction between P. eigii and P. pelasgis, both of which he noted shared acute hair tips and lack of callus hairs, as opposed to P. bulbosa (and P. hackelii lectotype). Figs 3 J, 3 K, 3 L, 10 A, 10 B, 10 C illustrate a normal habit, inflorescence and spikelet. Representative specimens. Turkey. Aydın (Gillespie et al. 10492 *, 10493 *). Antalya (Soreng et Cabi 9043 - a- 2 *). Bitlis (Tatvan, Kesan Deresi, 1800 m, Çelik 184 VANF). Erzurum (Uzundere, 2200–2400 m, A. Kaya 1244, p. p. b VANF). Izmir (Olfas 71 {c} E). Tekirdağ (Soreng et Cabi 9287). Uşak (?), (Geligan Dağ, entre Ala Cherr et Ouchak, Balansa s. n. pp. a G). Van (Murodiyea, 1750 m, Karaback K 1588 VANF). Croatia. (Flora Dalmatica, Cattaro, 17 May 1897, Baenitz s. n. [US- 04072277]). Greece. Karpathos (Poa pelasgis Type) and many other Aegean Islands, and Attica and Megarida, mapped and listed by Scholz (1985) and Strid (2016 a, 2016 b). Crete (Rechinger 13711 [US- 04072275]). Lesvos (Edmondson et McCintock 2376 E). Peloponnesos (Poa sinaica subsp. graeca Type)]. Cyprus. (7 Apr 2000, Scholz s. n. B, reported by H. Scholz, in Hand 2001). Israel. (Poa eigii Type HUJ; Simmons 4080 *, 4449, 4450 US; Bornmüller 1648 [US- 04072278]; Naftalsky HUJ 12596 [image!]; Zohary HUJ 40158 [image!]). Jordan. (Anonymous 1 [US- 04072276]; Dudin Farm, Semple 9 US; Hebron, Semple 31 US). Syria. (Hartmann [B 100594492]). Bulbous and non-bulbous plants of Poa subg. Poa sections Macropoa and Oreopoa reported or known from Turkey