Trachelophora lineata Aurivillius 1923

Trachelophora lineata Aurivillius, 1923 Fig. 2 A – E Trachelophora lineata Aurivillius, 1923: 465 (original description, type locality: Mount Penrissen, Borneo); Breuning 1950: 321 (key), 352; Heffern 2005: 28. Type material examined. Holotype: ♂ (NHRS), Trachelophora lineata Aur. (handwritten with black ink on a rectangular white label) / NHRS - ALJB 000000605 (printed with black ink on a rectangular white label) / 9471 E 92 + (printed with black ink on a rectangular blue label) / Penrissen May 1899 (handwritten and printed with black ink on a rectangular white label) / Type (printed with black ink on a rectangular white label) (Fig. 2 A – E). Redescription. (holotype, based on quality photographs, and modified from the original description) Male. Body length 17.0 mm, body width 5.5 mm. Head broad, slightly convex, densely and coarsely punctate; punctures umbilicate. Integument dark brown to blackish, largely obscured by dense recumbent yellowish-brown to ochraceous pubescence. Frons broad, weakly convex, densely covered with coarse, deep, round to slightly elongate punctures, irregularly arranged. Median groove broad and shallow, distinctly impressed on the frons, gradually fading toward both the clypeus anteriorly and the vertex posteriorly. Vertex broad, not elevated above the upper eye lobes, densely punctate as on frons, bearing a pair of conspicuous bright yellow longitudinal vittae extending from the upper margins of the antennal tubercles posteriorly. Genae short, moderately broad, weakly rounded, densely clothed with pale recumbent pubescence, not surpassing the outer margins of the eyes. Clypeus short, transverse, distinctly narrower than frons, densely punctate and pubescent, anterior margin straight to weakly rounded. Eyes large, coarsely faceted, strongly emarginate by antennal insertions; upper eye lobes narrow; lower eye lobes distinctly broader than the distance separating the upper lobes. Antennae slender, ~ 2 × the elytral length, reaching the elytral apex at antennomere VII. Scape clavate, slightly thickened apically, densely pubescent; remaining antennomeres cylindrical, clothed with short recumbent pubescence. Antennomeres X and XI subequal in length, each approximately equal to antennomere IX; antennomere III 4.0 mm long. Relative lengths of antennomeres (III = 1.00): scape 0.62, pedicel 0.10, IV 0.87, V 0.70, VI 0.62, VII – VIII 0.50, IX 0.45. Pronotum slightly wider than long, subquadrate; anterior and posterior margins nearly straight to weakly sinuate; lateral margins slightly rounded, each bearing a strong lateral spine. Disc coarsely punctate, with irregular vermiculate impressions; median region weakly elevated. Integument dark brown, densely covered with grayish- to yellowish-brown recumbent pubescence. Dorsum with five conspicuous bright yellow longitudinal vittae: one narrow median vitta and two pairs of lateral vittae extending continuously from the anterior to the posterior margin. Elytra elongate, subparallel-sided, gradually narrowed toward the apex. Surface densely and coarsely punctate, punctures deep and irregular, denser on the basal half; interspaces narrow and uneven, producing a strongly rugose appearance. Integument dark brown to blackish, irregularly mottled with yellowish pubescent maculae. Yellow markings not arranged into continuous longitudinal vittae, instead forming diffuse elongate patches along the lateral regions and smaller irregular maculae scattered over the basal, median, and apical thirds. Sutural region narrowly dark, nearly devoid of pale pubescence, contrasting distinctly with adjacent maculae. Pubescence short, dense, recumbent, partially concealing the integument. Legs long and slender. Femora clavate, gradually expanded toward the apex, densely clothed with grayish- to yellowish-brown pubescence, denser ventrally and apically. Tibiae more slender than femora, nearly straight with a weak median curvature, clothed with grayish to light-brown pubescence. Femora and tarsi bearing sparse white setae. Tarsi pseudotetramerous; metatarsomere I shorter than the combined length of metatarsomeres II and III; tarsomere III deeply bilobed, densely clothed with fine whitish setae. Pretarsal claws small, dark, weakly curved inward. Ventral surface densely clothed with yellowish-brown to pale ochraceous recumbent pubescence, forming irregular patches on the prosternum, mesoventrite, and metaventrite; prosternum, mesoventrite, and metaventrite bearing scattered yellowish setae. Prosternal process narrow and elongate, extending between the procoxae; procoxal cavities open posteriorly. Hypomera densely clothed with adpressed pubescence. Abdominal ventrites I – V densely clothed with yellowish-brown pubescence; ventrites II – IV each with distinct pale pubescent patches; ventrite I longest. Interventral sutures straight and distinct. Ventrite V gradually narrowed posteriorly, broadly rounded apically, densely pubescent, with scattered longer setae along the apical margin. Remarks. The combination of paired yellow vertex stripes, five-striped pronotum, irregular elytral maculation, and dense ochraceous pubescence readily distinguishes this species from all congeners. This species was described by Aurivillius (1923), with the type locality [Mount] Penrissen, Borneo, and the type specimen deposited in NHRS. The other specimen deposited in the NMB was misidentified by Breuning as T. lineata, but in fact represents a female of T. curvicollis (see comments under T. curvicollis). Trachelophora lineata and T. curvicollis share the presence of a yellow vitta extending from the pronotum to the basal part of the elytra, as well as similar coloration of the pubescence covering the elytra. However, they differ markedly in several diagnostic characters. In T. lineata, the frons bears a shallow, short median impression, which becomes obsolete toward the clypeus, whereas in T. curvicollis the frons exhibits a distinct, deep median groove extending nearly to the clypeus. The pronotum of T. lineata is wider than long and subquadrate, while in T. curvicollis the pronotum is longer than wide. The elytral apex of T. lineata is strongly obliquely truncate, with the outer apical angle produced into a long, sharp spine, whereas in T. curvicollis the elytral apex is obliquely truncate, with the outer apical angle only shortly produced. Distribution. Mt. Penrissen, Borneo.