Xenophrys vuquangensis sp. nov. (Anura, Megophryidae), a new horned frog species from Ha Tinh Province, Vietnam

A new species of Xenophrys is described from Vu Quang National Park, Ha Tinh Province, Vietnam based on morphological and molecular differences. Morphologically, Xenophrys vuquangensis sp. nov. is distinguishable from its congeners by a combination of the following morphological characteristics: size medium (SVL 63.9–68.6 mm in males, 77.6 mm in the female); tympanum distinct; vomerine teeth present; tongue feebly notched posteriorly; external vocal sac present; toes with rudimentary webbing; subarticular tubercles absent; lateral fringes narrow along toes; dorsal skin of body with small granules; flank with numerous large tubercles; dorsum with a dorsal ridge, in >-< shape; belly and ventral surface of thigh with numerous tubercles; dorsal surface of the head and back reddish brown with a dark brown triangle between the eyes; a dark brown marking edged, in >-< shape, along dorsal ridge; and flank and back of thigh with larger white spots. In the phylogenetic analyses, the new species is nested within the genus Xenophrys and placed as a sister taxon to X. truongsonensis with interspecific uncorrected genetic p-distances, based on the mitochondrial 16S rRNA gene, ranging from 4.0% (compared with Xenophrys maosonensis) to 14.8% (compared with Xenophrys awuh).