Composetia marmorata

Composetia marmorata (Horst, 1924) Fig. 5 A – D Nereis (Ceratonereis) marmorata Horst, 1924: 177–178, pl. 34, figs 13–16. Composetia marmorata. – Pamungkas and Glasby 2015: 8 –9, fig. 3 C, D. Type locality. Between Geser and Seram Islands, Maluku Province, Indonesia. Material examined. MZB.Pol.00314 (7 males) and MZB.Pol.00315 (1 female), collected as heteronereids locally known as ‘ nyale ’ by Dewi C. Murniati on 8 March 2018 from Lombok Island: Seger Beach, Kuta Village, Pujut District, Central Lombok Regency, West Nusa Tenggara Province (8°54'25.9"S, 116°17'49.3"E); SMF 33463 (3 specimens), collection details same as MZB.Pol.00314; MZB.Pol.00276 (13 specimens), collected by Joko Pamungkas on 23 November 2017 from Warmar Island: Siwalima Village, Aru Islands District, Aru Islands Regency, Maluku Province (5°45'26.9"S, 134°13'23.5"E); MZB.Pol.00277 (1 specimen), collected by Joko Pamungkas on 21 November 2017 from Ambon Island: Hutumuri Village, South Leitimur District, Ambon City, Maluku Province (3°41'46.1"S, 128°17'49.0"E); MZB.Pol.00339 (2 specimens), collected by Joko Pamungkas on 24 November 2017 from Warmar Island: Wangel Village, Aru Islands District, Aru Islands Regency, Maluku Province (5°45'05.1"S, 134°11'43.2"E). Comparative material. Composetia marmorata as heteronereids locally known as ‘ laor / wawo ’. MZB.Pol.00161 (3 males), MZB.Pol.00162 (2 females), RCLA.Ann.042 (3 males), RCLA.Ann.043 (1 female), RCLA.Ann.044 (3 males), RCLA.Ann.045 (2 females), collected by Robert Alik on 18–19 March 2014 from Ambon Island: Suli Village, Salahutu District, Central Maluku Regency, Maluku Province (3°37'38.2"S, 128°18'25.0"E); RCLA.Ann.046 (1 male), collected by Abraham S. Leatemia on 19 March 2014 from Ambon Island: Mahia Subvillage, Urimessing Village, Nusaniwe District, Ambon City, Maluku Province (3°44'42.6"S, 128°11'24.6"E). Diagnosis. Body color in alcohol cream with distinctive marmorated stripes on prostomium and anterior body. All paragnaths conical, arranged as follows. Area I = 1–2; Area II = 7–14; Area III = 9–13; Area IV = 14–16; Areas V – VIII absent. Male and female epitokes morphologically similar, yet males with sub-distally swollen (not scalloped) dorsal cirri in natatory region (unmodified in female) and pygidial rosette (lacking in females). Pre-natatory region comprising 17 or 18 chaetigers with basally swollen dorsal cirri on chaetigers 1–6 or 7, and basally swollen ventral cirri on chaetigers 1–5. Natatory region extending to pygidium. Females with unpigmented eggs (Pamungkas and Glasby 2015). Distribution and habitat. The species was found in Maluku and West Nusa Tenggara (Indonesia) and Lizard Island (NE Australia), inhabiting intertidal rocky shore. Remarks. Although the distinctive marmorated (= veined) stripes of Composetia marmorata appear to be faded on the present specimens, the paragnath counts match those described by Pamungkas and Glasby (2015) for Ambonese specimens and Horst’s (1924) original description. In addition, the account of Composetia marmorata swarming as nyale on Lombok Island agrees well with the description of the species swarming as laor / wawo on Ambon Island. The non-reproductive stage specimens obtained in the present work, as well as nyale, complement the collection of this species at MZB.