Dodona dipoea subsp. astroides Hu, L. - P. Zhang & Inayoshi 2026, ssp. nov.

Dodona dipoea astroides Hu, L.-P. Zhang & Inayoshi ssp. nov. [ĒaeDzợāē亚ª] Description: Male (Figure 6A–D): Forewing length 18–20 mm (mean = 18.6 ± 0.5 mm, n = 7). Forewing upperside dark brown, with rows of scattered greyish-ochreous spots: sub-basal ochreous band across cell CuA 2 and the discocell; series of discal spots in cells CuA 2 (paired), CuA 1 (oblique), M 2 and M 1 (obliquely paired) and costal spots (small obliquely paired); series of postdiscal spots in cells 2A and CuA 2 (linear), CuA 1 (single, dark), M 3 (large and small), M 2 (single), M 1 (single), costal (paired) and subapical (double) forming forked pattern; no submarginal spots. Forewing underside pale chocolate brown, lower half darker, with markings reflecting upperside but paler, more pronounced, expanded and clearly defined, thin obscure transverse band at end of discoidal cell. Hindwing upperside dark ochreous brown with obscure discal band; postdiscal dark ochreous band across cells M 2, M 1, and R 5; obscure double submarginal ochreous-grey bands connecting to two black ocelli in cells M 1 and R 5; tornal lobe black. Hindwing underside pale chocolate brown with clear bands and spots: two straw-coloured basal to tornal bands; broad sub-basal band from costa (white) to middle of cell CuA 2 (turned straw-coloured); white costal spot lined with black; thin and faint discal whitish band lined with black in CuA 2 and CuA 1 connecting to tornal markings; white postdiscal band lined with black in cells M 3 to M 1; obscure double submarginal bands connecting to two black ocelli with white edges in cells M 1 and R 5; tornal lobe black crowned by white lines and grey patch. Female (Figure 6E): Colour and markings similar to male, both wings with more convex termen. Male genitalia (Figure 7A): Moderate to heavily sclerotized. Vinculum slender, posteriorly slightly curved near tegumen; uncus hook-beak like in lateral view and hoody like in posterior view, tip acute and down curved, both dorsal and lateral surfaces covered with setae; brachium strong with pointed tip; saccus short. Valves and juxta merged, carina penis connected with basal 3/4 portion of phallus, tip deeply bifid; base of valve broad and narrowed toward tip, tip of valve roundish with deeply concave and acute point, ventral and dorsal margins thick covered with long setae, tip of ventral margin bluntly projected; harpe absent. Juxta tongue-shaped. Phallus tubular, long, robust and curved in median portion, tip blunt, cornutus mostly membranous. Female genitalia (Figure 7B): Papillae anales semi-circular, margin covered with short setae; apophyses posteriores weak. Ostial opening partially covered by sclerotized lamella postvaginalis slightly concave on ventral view, lamella antevaginalis membranous. Ductus bursae long, median portion with sclerotized longitudinal striae. Corpus bursae oval-bulbous, membranous with two roundish and spiny signa patches, one near base, one near end. Etymology: The subspecies name reflects its morphological resemblance with D. astra sp. nov. sympatric in northwestern Yunnan. Type materials: HOLOTYPE: ♂, Renqingbeng Monastery, Motuo County, Nyingtri Region, Tibet, China, 2025-V-25, Shao-Mian Zhang et al. leg. [SWI]. PARATYPES: 1♂, Gelin, Motuo County, Nyingtri Region, Tibet, China, 2026-VI-26, Rui-Qi Hu leg. [YNU]. 3♂♂, Road from Dulongjiang to Cikai, Gongshan County, Nujiang Prefecture, Yunnan, China; 2023-VI-28; Hui-Hong Zhang leg. [YNU]. 1♂, northeastern Putao, Kachin State, Myanmar; 2011-VI-25; Prasobsuk Sukkit leg. [YI]. 1♂, same locality; 2003-V-30; Prasobsuk Sukkit leg. [YI]. 2♂♂, same locality; 1996-VI-19; Prasobsuk Sukkit leg. [YI]. 1♀, the same locality; 2011-V-27; Prasobsuk Sukkit leg. [YI]. Distribution: The new subspecies is currently found in Gongshan and Dulongjiang areas of northwestern Yunnan, China, and also recorded from Motuo, southern Tibet, China and Kachin State, Myanmar. Habitat and behaviour: Like the nominotypical subspecies in Nepal and ssp. dipoeides in Shangri-La, this species requires forest habitat but is more adapted to anthropogenic disturbance (Figure 8). Males fly near the forest edge or along paved roads near the forest. Territorial behaviour on shrubs, trees or rocks and mud puddling were observed.