Coexistence, Oscillations, and Chaos in a Discrete Predator-Prey System with Fear-Modulated Prey Growth

Predator–prey interactions are governed not only by direct consumption but also by behavioral responses of prey to the perceived risk of predation. In this work, we formulate a discrete-time predator–prey model in which predator-induced fear modifies prey reproduction through an exponential suppression mechanism, while predator consumption follows a Holling type-II functional response. The resulting map combines Ricker-type prey growth, density-dependent predation, and a non-consumptive effect associated with predator presence. We establish fundamental dynamical properties of the system by proving positivity, boundedness, and persistence under appropriate parameter restrictions. The existence of biologically meaningful equilibria is determined, and their local behavior is characterized using the Jacobian matrix and Jury stability criteria. To examine the influence of ecological parameters on the dynamics, we employ a global sensitivity analysis based on the Partial Rank Correlation Coefficient (PRCC) approach. The analysis identifies the parameters that most strongly affect prey and predator abundance. Numerical investigations reveal a wide range of dynamical regimes, including stable coexistence, periodic oscillations, higher-period attractors, quasi-periodic motion, and chaos. In particular, flip and Neimark–Sacker bifurcations are observed as ecological parameters vary. Basin-of-attraction computations and two-parameter iso-spike diagrams further demonstrate the presence of multistability and strong dependence on parameter combinations and initial conditions. Finally, period-doubling control and pole-placement techniques are employed to suppress undesirable chaotic oscillations and recover stable coexistence. The results emphasize that predator-induced fear can substantially modify population fluctuations and may act as an important mechanism regulating coexistence and complex dynamics in discrete ecological systems.

Authors

Institutions

Publication Details

Journal
Mathematics
Published
2026-09-16
DOI
https://doi.org/10.3390/math14183371
Primary Topic
Mathematical and Theoretical Epidemiology and Ecology Models
Type
article
Field-Weighted Citation Impact
0.00
Controls
|||
ALL TIME
JAN
FEB
MAR
APR
MAY
JUN
JUL
AUG
SEP
article

Coexistence, Oscillations, and Chaos in a Discrete Predator-Prey System with Fear-Modulated Prey Growth

Sk. Sarif Hassan, A. M. Ełaiw, N. H. AlShamrani, Sujay Goldar et al.
Mathematics
Mathematical and Theoretical Epidemiology and Ecology Models
article

Coexistence, Oscillations, and Chaos in a Discrete Predator-Prey System with Fear-Modulated Prey Growth

Sk. Sarif Hassan, A. M. Ełaiw, N. H. AlShamrani, Sujay Goldar, A. Mohsen, Purnendu Sardar
article en

Abstract

Predator–prey interactions are governed not only by direct consumption but also by behavioral responses of prey to the perceived risk of predation. In this work, we formulate a discrete-time predator–prey model in which predator-induced fear modifies prey reproduction through an exponential suppression mechanism, while predator consumption follows a Holling type-II functional response. The resulting map combines Ricker-type prey growth, density-dependent predation, and a non-consumptive effect associated with predator presence. We establish fundamental dynamical properties of the system by proving positivity, boundedness, and persistence under appropriate parameter restrictions. The existence of biologically meaningful equilibria is determined, and their local behavior is characterized using the Jacobian matrix and Jury stability criteria. To examine the influence of ecological parameters on the dynamics, we employ a global sensitivity analysis based on the Partial Rank Correlation Coefficient (PRCC) approach. The analysis identifies the parameters that most strongly affect prey and predator abundance. Numerical investigations reveal a wide range of dynamical regimes, including stable coexistence, periodic oscillations, higher-period attractors, quasi-periodic motion, and chaos. In particular, flip and Neimark–Sacker bifurcations are observed as ecological parameters vary. Basin-of-attraction computations and two-parameter iso-spike diagrams further demonstrate the presence of multistability and strong dependence on parameter combinations and initial conditions. Finally, period-doubling control and pole-placement techniques are employed to suppress undesirable chaotic oscillations and recover stable coexistence. The results emphasize that predator-induced fear can substantially modify population fluctuations and may act as an important mechanism regulating coexistence and complex dynamics in discrete ecological systems.

MathematicsVol. 14(18)
University of Baghdad (IQ), Jadavpur University (IN), King Abdulaziz University (SA), Kanya Maha Vidyalaya (IN), University of Jeddah (SA)
Openalex Percentile: Top 8%
Mathematical and Theoretical Epidemiology and Ecology Models
AI Navigator

Ask Laika to Summarize, Analyze, and Connect papers live on the map.

Summarize Papers & Methodologies

Extract key findings, datasets, and comparative methods across publications.

Benchmark Rankings & Visual Analytics

Rank top research institutions, authors, funders, topics, and journals by Field-Weighted Citation Impact (FWCI) and paper volume with instant charts.

Connect Distant Disciplines

Bridge topological clusters on the map to find hidden collaborative intersections.