Spatial Patterns of Soil Water-Holding Capacity and Their Environmental Drivers in Spruce-Fir-Korean Pine Forest of the Xiaoxing’an Mountains

In the primary spruce-fir-Korean pine forest affected by historical windthrow, soil water-holding capacity shows complex spatial associations with forest microenvironment and soil physicochemical properties. However, its spatial variability and multi-factor hierarchical association pathways remain poorly understood. Taking the primary spruce-fir-Korean pine forest on a historically windthrow-affected site in the Liangshui National Nature Reserve, Xiaoxing’an Mountains, China as the research object, geostatistics and partial least-squares structural equation modeling were used to analyze the spatial pattern of topsoil (0–20 cm) water-holding capacity and its environmental association pathways. The results showed saturated, capillary and field water-holding capacities of topsoil exhibited moderate variability and strong spatial autocorrelation (all nugget to sill ratios < 25%), showing a patchy distribution. No significant direct association was detected between windthrow mechanical disturbance intensity and topsoil water-holding capacity. Bulk density (path coefficient = −0.685, p < 0.001) and soil porosity (path coefficient = 0.273, p < 0.001) had significant direct associations with soil water-holding capacity, whereas soil particle size distribution showed no significant effect (p > 0.05). Canopy structure and understory microclimate exerted indirect associations with soil water-holding capacity via soil structure; litter showed no significant associative effect (p > 0.05). These results indicate that soil water-holding capacity on the historically windthrow-affected site was not randomly distributed, but presented an ordered patchy pattern closely related to in-plot micro-environmental factors.