Post-Collapse Complexity Cascade (PCCC v2.3): A Phenomenological Model of Post-Extinction Macroevolutionary Recovery Dynamics

Background: The Modern Synthesis explains adaptive evolution under relatively stable environmental conditions but provides limited mechanistic explanation for rapid post-extinction radiations and punctuated evolutionary transitions. Major biological reorganizations following catastrophic events suggest that non-equilibrium environmental conditions may accelerate the emergence of novel adaptive configurations. Model: We formalize a revised Post-Collapse Complexity Cascade (PCCC) framework in which mass extinction events function as high-entropy perturbations Ω(Ω₀) that generate transient far-from-equilibrium environments. Under these conditions, persistent molecular substrates — including cryptobiotic structures, lipid assemblies, prion-like conformational templates, and dormant genetic variation — undergo accelerated reorganization. Core Equation:dΨ/dt = k · R(t) · ∇μ(t) − δΨR(t) = Σ αᵢ Iᵢ e^(−λᵢ(t−t₀)) Where:Ψ = emergence rate of novel configurationsR(t) = persistence-weighted reservoir∇μ(t) = thermodynamic gradient Results: The model predicts:1. Lag times 10–30% between collapse and innovation2. Wave-like emergence of new complexity3. Reuse of pre-existing dormant variation Predictions match events from Great Oxidation Event (2.4 Gya) → Toba supereruption (74 kya). Case study: human evolution reorganization waves 50–8 kya, admixture patterns & cultural transitions. Conclusions: PCCC extends evolutionary theory by providing measurable thermodynamic mechanism for rapid innovation. Falsifiable via ancient DNA analysis; fully compatible with Extended Evolutionary Synthesis. ✅ Version 2.1 | Final Lengkap✅ Preprint Resmi | Belum melalui peer-review✅ DOI: 10.5281/zenodo.20129xxx✅ Arsip riwayat:• Awal: 10.5281/zenodo.20090501• V1: 10.5281/zenodo.19637169• V2: 10.5281/zenodo.20129237 ✅ Version 2.3.1 | Koreksi Alamat Penulis: Bojonegoro, Jawa Timur✅ Ditambahkan ORCID: 0009-0006-9815-982X✅ DOI Utama: 10.5281/zenodo.20129237