Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye

Among Paleozoic foraminifers, miliolids represent one of the most diverse groups, which reached their highest diversity during the Middle to Late Permian. Notably, many taxa within the family Neodiscidae emerged during the Guadalupian epoch. In this study, two fossiliferous Middle Permian blocks were studied for their foraminiferal assemblages around Kutludüğün Yayla (east of Ankara, Türkiye) from the Elmadağ Olistostrome. We present a systematic and phylogenetic revision of the subfamilies Neodiscinae and Agathammininae, and propose the erection of Globidiscinae subfam. nov. included within the family Neodiscidae. The foraminiferal genera Kamurana, Globidiscus, and Agathammina are taxonomically re-evaluated and emended based on newly described miliolid taxa, in conjunction with a reassessment of previously published data. The composition and taxonomic status of the subfamily Neodiscinae is revised. The subfamily Agathammininae and family Neodiscidae are emended. The genus Neodiscus derives from Praeneodiscus in the Capitanian, and Kamurana, Globidiscus, Rectogordiopsis gen. nov., Davanela gen. nov. and Kaganella gen. nov. derived probably from Neodiscus in the late Capitanian. The new taxa are Globiscinae subfam. nov.; Rectogordiopsis gen. nov.; Davanella gen. nov. and Kaganella gen. nov., Rectogordiopsis kamuranaeformis gen. et sp. nov., Rectogordiopsis ovaliformis gen. et sp. nov., Davanella ankaraensis gen. et sp. nov., Davanella acuminata gen. et sp. nov., Kaganella tekini gen. et sp. nov.