Integrative taxonomic revision of Capilla Grishin, 2023, subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera, Hesperiidae, Pyrginae, Carcharodini), with descriptions of four new species

The taxonomy of the subgenus Staphylus (Capilla) is reviewed, including redescriptions of known species, identification keys, and detailed distribution maps. The female genitalia of Staphylus (Capilla) azteca (Scudder, 1872), S. (C.) caribbea (Williams & Bell, 1940), S. (C.) corumba (Williams & Bell, 1940), S. (C.) eryx Evans, 1953, S. (C.) tucumanus (Plötz, 1884) and S. (C.) tyro (Mabille, 1878) are described and illustrated for the first time. Four new species are described: Staphylus (Capilla) nicoleae Lemes sp. nov . from Colombia and Venezuela, S. (C.) ricardoi Lemes, sp. nov . from Peru, S. (C.) neideae Lemes sp. nov . and S. (C.) neivae sp. nov . from Brazil. Pholisora imperspicua Hayward, 1940 is a syn. nov . of Staphylus ( C.) lizeri lizeri (Hayward, 1938) and Hesperia melangon epicaste Mabille, 1903 is a syn. nov . of Staphylus ( C.) melangon melangon (Mabille, 1883). Neotypes are designated for Staphylus epicaste Mabille, 1903, Nisoniades tucumanus Plötz, 1884 and Staphylus fascia Hayward, 1933. Lectotypes are designated for Pholisora azteca Scudder, 1872, Bolla machuca Schaus, 1913, Helias tyro Mabille, 1878, Staphylus anginus Schaus, 1902, Hesperia melangon 1883, Hesperia musculus Burmeister, 1875 and Helias aurocapilla Staudinger, 1876.