<i>Orangutan</i> : An R Package for Analyzing and Visualizing Phenotypic Data in the Context of Species Descriptions and Population Comparisons

Phenotypic characters have long been central to species diagnosis and remain indispensable even in the age of genomics. However, phenotypic datasets are often complex-spanning dozens of traits of varying types and units, with correlated variables and unbalanced sampling-posing challenges for robust, reproducible analysis. Existing software solutions are fragmented, usually requiring labor-intensive workflows across multiple tools and manual steps, which undermines reproducibility and hinders comparisons across studies. To address these methodological and practical challenges, I introduce <i>Orangutan</i>, an R package designed to provide a reproducible, easy-to-implement framework for comparing groups using mensural and meristic data. <i>Orangutan</i> integrates statistical analysis and visualization for species diagnosis and population comparisons within a single workflow. The package streamlines the identification of diagnostic, nonoverlapping traits between species, while enabling rigorous assessment of both individual and multivariate trait differences in overlapping traits. Core features include optional allometric correction to remove size effects, optional outlier removal, automated selection of appropriate univariate tests with post hoc comparisons, and integrated multivariate analyses. All outputs, including tables and publication-ready figures, are generated with minimal coding, ensuring accessibility and standardization. Empirical validation with real-world datasets-including animal and plant species-demonstrates that <i>Orangutan</i> robustly identifies diagnostic traits, reveals both subtle and clear group differences, and achieves high classification accuracy with phenotypic data alone. By automating and unifying key analytical steps, <i>Orangutan</i> promotes reproducibility, transparency, and efficiency in phenotypic research. This package could empower researchers in taxonomy, ecology, and evolutionary biology to adopt quantitative good practices for species diagnoses, facilitating comparative studies and advancing methodological standards in morphological data analysis. <i>Orangutan</i> is freely available as open-source software with comprehensive documentation to facilitate broad adoption.

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Publication Details

Journal
Ecology and Evolution
Published
2026-02-01
DOI
https://doi.org/10.1002/ece3.73111
Primary Topic
Genetic diversity and population structure
Type
article
Field-Weighted Citation Impact
0.00

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article

Orangutan : An R Package for Analyzing and Visualizing Phenotypic Data in the Context of Species Descriptions and Population Comparisons

Javier Torres
Ecology and Evolution
Genetic diversity and population structure
article

Orangutan : An R Package for Analyzing and Visualizing Phenotypic Data in the Context of Species Descriptions and Population Comparisons

Javier Torres
article en

Abstract

Phenotypic characters have long been central to species diagnosis and remain indispensable even in the age of genomics. However, phenotypic datasets are often complex-spanning dozens of traits of varying types and units, with correlated variables and unbalanced sampling-posing challenges for robust, reproducible analysis. Existing software solutions are fragmented, usually requiring labor-intensive workflows across multiple tools and manual steps, which undermines reproducibility and hinders comparisons across studies. To address these methodological and practical challenges, I introduce <i>Orangutan</i>, an R package designed to provide a reproducible, easy-to-implement framework for comparing groups using mensural and meristic data. <i>Orangutan</i> integrates statistical analysis and visualization for species diagnosis and population comparisons within a single workflow. The package streamlines the identification of diagnostic, nonoverlapping traits between species, while enabling rigorous assessment of both individual and multivariate trait differences in overlapping traits. Core features include optional allometric correction to remove size effects, optional outlier removal, automated selection of appropriate univariate tests with post hoc comparisons, and integrated multivariate analyses. All outputs, including tables and publication-ready figures, are generated with minimal coding, ensuring accessibility and standardization. Empirical validation with real-world datasets-including animal and plant species-demonstrates that <i>Orangutan</i> robustly identifies diagnostic traits, reveals both subtle and clear group differences, and achieves high classification accuracy with phenotypic data alone. By automating and unifying key analytical steps, <i>Orangutan</i> promotes reproducibility, transparency, and efficiency in phenotypic research. This package could empower researchers in taxonomy, ecology, and evolutionary biology to adopt quantitative good practices for species diagnoses, facilitating comparative studies and advancing methodological standards in morphological data analysis. <i>Orangutan</i> is freely available as open-source software with comprehensive documentation to facilitate broad adoption.

Ecology and EvolutionVol. 16(2)
University of Nebraska–Lincoln (US)
University of Nebraska-Lincoln
Openalex Percentile: Top 46%
Genetic diversity and population structure
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