Late Devonian Ichnofossil Lagerstätte of the Eastern Anti-Atlas, Morocco

A rich ichno-assemblage is known from the Upper Devonian (Famennian) Aoufilal Formation of the eastern Anti-Atlas in Morocco. Despite its high diversity, only a few trace fossils have been properly described until now. In addition to fish-produced ichnotaxon Undichna detected earlier, resting and walking trace fossils of aquatic arthropods (Rusophycus and Diplichnites), walking trackways and feeding areas of eurypterids (Palmichnium), and rosette-type crustacean or polychaetan feeding burrows (Asterichnus) are described here. All the traces are preserved in positive hyporeliefs on the lower bedding surfaces of a reddish-brown fine-grained sandstone accumulated under shallow-marine conditions. A high diversity, excellent preservation and frozen state of the paleocommunity despite the absence of body fossils allow us to rate the Aoufilal ichno-assemblage as an ichnofossil Lagerstätte.