Paleosols, Floristic Paleosuccession, and a New Genus of Fossil Tetrapod Traces from the Upper Permian of the North of European Russia

This paper discusses the possible depositional settings of the Upper Permian beds (Severodvinian and Vyatkian stages), exposed in the basin of the Sukhona and Malaya Severnaya Dvina rivers. Carbonate paleosols and floristic paleosuccession that took place during the Late Permian epoch in this region are considered. A new ichnogenus and a new ichnospecies (Gorgonopus opokensis Naugolnykh Ichnogenus et Ichnospecies nov.) are described.