Discrepant Pathway in Regulating ET Under Change in Community Composition of Alpine Grassland in the Source Region of the Yellow River

Understanding evapotranspiration (ET) dynamics under community composition transitions in grasslands is crucial for interpreting alpine ecosystem responses to climate change. We investigated variations in ET and its components during the growing season across five alpine grassland transition types in the Source Region of the Yellow River (SRYR) from 1986 to 2018, integrating climatic, vegetation, and soil factors. Under warming and wetting conditions, ET increased significantly by 1.17 mm yr−1, accounting for 79.39% of annual precipitation, while soil moisture declined slightly. A pronounced temperature–precipitation decoupling emerged between alpine meadow-origin (AM-origin) and alpine steppe-origin (AS-origin) transitions, indicating differential hydrological responses driven by community composition. Vegetation growth increased across all transitions, yet its regulation of ET components varied by transition type. Transpiration dominated ET increases, contributing over 80% in AM-origin and 100% in AS-origin transitions. Soil evaporation exhibited contrasting trends: decreasing in AS-origin transitions due to enhanced soil insulation from vegetation growth, but increasing in AM-origin transitions, thereby reducing soil moisture. Interannual ET growth rates and seasonal fluctuations were greater in AM-origin than in AS-origin transitions. A critical turning point in ET trends, caused by changes in precipitation, revealed the divergent hydrological trajectories among the transitions. In AM-origin transitions, temperature primarily drove ET increases, causing soil drying (strongest in AM to TS), whereas in AS-origin transitions, precipitation dominated, resulting in soil wetting (more pronounced in AS to AM). These findings demonstrate that the directionality of compositional transitions governs hydrological responses more strongly than absolute vegetation states.