Haploniscus bulbosus Henseler, Knauber & Riehl 2025, sp. nov.

Haploniscus bulbosus Henseler, Knauber & Riehl sp. nov. Materials Type status: Holotype. Occurrence: catalogNumber: SMF 62946; recordNumber: KBHap 003; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: C7CBAC4D-BD5E-557E-B66D-85C1D2AF559A; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: collectionID: Crustacea; institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: SMF 62947; recordNumber: KBHap 004; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: 1F5497CF-228B-5015-B6B1-293994CF9A30; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: collectionID: Crustacea; institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62948; recordNumber: KBHap 005; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: 297F6FA9-A715-5427-BD57-EBCC4CEDFAAB; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62949; recordNumber: KBHap 011; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: C3ADCEA9-307F-5AF8-86FB-BF41665F52AD; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 23. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-09 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62950; recordNumber: KBHap 012; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: DE20034C-9543-50F7-BE92-8F2982423588; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 23. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-09 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62951; recordNumber: KBHap 013; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: DDC1F330-B393-54FF-B6BD-3E96D6A23ED4; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to t

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Zenodo (CERN European Organization for Nuclear Research)
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2025-10-15
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https://doi.org/10.5281/zenodo.17361942
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Data Analysis with R
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article

Haploniscus bulbosus Henseler, Knauber & Riehl 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Data Analysis with R
article

Haploniscus bulbosus Henseler, Knauber & Riehl 2025, sp. nov.

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Abstract

Haploniscus bulbosus Henseler, Knauber & Riehl sp. nov. Materials Type status: Holotype. Occurrence: catalogNumber: SMF 62946; recordNumber: KBHap 003; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: C7CBAC4D-BD5E-557E-B66D-85C1D2AF559A; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: collectionID: Crustacea; institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: SMF 62947; recordNumber: KBHap 004; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: 1F5497CF-228B-5015-B6B1-293994CF9A30; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: collectionID: Crustacea; institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62948; recordNumber: KBHap 005; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: 297F6FA9-A715-5427-BD57-EBCC4CEDFAAB; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean KuramBio station SO 223-10 - 09; verbatimDepth: 5264-5266 m; verbatimLatitude: 41°11'16.188''N; verbatimLongitude: 150°5'36.888''E; decimalLatitude: 41.18783; decimalLongitude: 150.09358; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 26. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-10 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62949; recordNumber: KBHap 011; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: C3ADCEA9-307F-5AF8-86FB-BF41665F52AD; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 23. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-09 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62950; recordNumber: KBHap 012; individualCount: 1; sex: female; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: DE20034C-9543-50F7-BE92-8F2982423588; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to the Kurile Kamchatka Trench and abyssal plain on board of the R / V SONNE, 223 rd Expedition, 21 July - 7 September 2012. University of Hamburg; Biozentrum Grindel & Zoologisches Museum Hamburg, Hamburg. Cruise Report, 100 pp. Available from: http://edok01.tib.uni-hannover.de/edoks/e01fb13/741102293.pdf.; eventDate: 23. 08. 2012; habitat: abyssal sediment; fieldNumber: SO 223-09 - 09; Record Level: institutionCode: SMF; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: SMF 62951; recordNumber: KBHap 013; individualCount: 1; sex: male; lifeStage: adult; preparations: whole animal (ETOH); occurrenceID: DDC1F330-B393-54FF-B6BD-3E96D6A23ED4; Taxon: scientificName: Haploniscus bulbosus Henseler, Knauber & Riehl; kingdom: Animalia; phylum: Arthropoda; class: Malacostraca; order: Isopoda; family: Haploniscidae; genus: Haploniscus; specificEpithet: bulbosus; scientificNameAuthorship: Henseler, Knauber & Riehl; nomenclaturalCode: ICZN; Location: higherGeography: Pacific Ocean; locality: North-west Pacific Ocean, KuramBio station SO 223-09 - 09; verbatimDepth: 5399-5421 m; verbatimLatitude: 40°34'22.8''N; verbatimLongitude: 150°59'54.888''E; decimalLatitude: 40.573; decimalLongitude: 150.99858; Identification: identifiedBy: Henry Knauber; Event: samplingProtocol: Benthos trawl, Camera-Epibenthic Sledge, sieved through 0.3 mm mesh | Riehl T, Brenke N, Brix S, Driskell A, Kaiser S, Brandt A (2014) Field and laboratory methods for DNA studies on deep-sea isopod crustaceans. Polish Polar Research 35: 205–226. https://doi.org/10.2478/popore-2014-0018 | Brandt A, Malyutina MV (2012) The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Study): to t

Zenodo (CERN European Organization for Nuclear Research)
Goethe University Frankfurt (DE), Hofstra University (US), Hong Kong Baptist University (HK), Scripps Institution of Oceanography (US), Universidade Estadual de Campinas (UNICAMP) (BR), Chinese Academy of Sciences (CN), Universidade Federal de Pernambuco (BR), Zoological Institute (RU), Senckenberg Research Institute and Natural History Museum Frankfurt/M (DE), University of Łódź (PL), Institute of Oceanology (CN), Muséum national d'Histoire naturelle (FR), Universidad Nacional Autónoma de México (MX)
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