Euspira incus Harzhauser, Landau & Guzhov, 2025, nov. sp.

Euspira incus nov. sp. Figs 2N, 37A 1 –A 4 Type material. Holotype: NHMW 2024 /0240/0002, SL: 17.3 mm, MD: 18.3 mm, Lăpugiu de Sus (Romania), Figs 2N, 37A 1 –A 4. Type locality. Lăpugiu de Sus (Romania), Făget Basin. Type stratum. Silt and clay of the Dej Formation. Age. Middle Miocene, Badenian (Langhian). Etymology. Incus, Latin for anvil, referring to the knob-like umbilical callus; noun in apposition. Diagnosis. Small, depressed globular shell with low conical spire, strongly expanding last whorl, narrow umbilical groove with knob-like umbilical callus and prominent anterior lobe of parietal callus. Description. Small, depressed globular shell of four teleoconch whorls; slightly wider than high (SL/MD = ~0.9); spire low conical of strongly convex whorls, apical angle 125°. Protoconch poorly preserved, low turbiniform of about two convex whorls. Suture linear, slightly impressed. Teleoconch whorls with indistinct subsutural ramp, poorly delimited by widely rounded shoulder. Periphery and base strongly convex. Last whorl strongly inflated, attaining ~90% of total height. Periphery below mid-whorl (~44%). Growth lines delicate and weakly prosocyrt in apical view, subobsolete, prosocline at periphery. Aperture D-shaped, prosocline, attaining ~80% of total height. Position of adapical tip of aperture high (~87%), slightly below shoulder. Opercular ride weak. Columellar lip strongly reinforced, basal lip and outer lip not preserved. Columellar angle ~28°. Parietal callus long (~58%), strongly thickened with prominent adapical swelling and deeply concave margin; not expanding over base. Anterior lobe moderately thick, forming short tongue. Umbilicus narrow (~9%), moderately deep, reduced to semicircular groove; funicle not visible. Umbilical callus knob-like, semicircular, separated from anterior lobe by concavity. Basal fasciole solid, weakly angled towards aperture. Discussion. This species is characterized by its knob-like umbilical callus delimited by a relatively narrow umbilical groove. Euspira umbilicocarinata Robba, Pedriali & Quaggiotto, 2016, from the Late Miocene of Italy, is reminiscent of Euspira incus nov. sp. but has a smaller umbilical knob (see Robba et al. 2016: pl. 10, fig. 5). Note that an opercular ridge can be present in Polinicinae although their operculum is not calcified. Paleoenvironment. Unknown. Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Lăpugiu de Sus (Romania) (hoc opus).