Hanleyidae Bergenhayn 1955

Family Hanleyidae Bergenhayn, 1955 Genus Hanleya Gray, 1857 Type species. Hanleya debilis Gray, 1857 (= Chiton hanleyi Bean in Thorpe, 1844), by monotypy. ......continued on the next page ......continued on the next page Distribution. The genus Hanleya is recorded from the lower Oligocene onwards. Currently, all known recent species of Hanleya occur only in the Atlantic Ocean and adjacent seas, from off Brazil (25.44° S) to the Barents Sea (74.27° N) (Sirenko et al. 2016). The fossil record includes the Oligocene of Germany (Janssen 1978), the Middle Miocene of Paratethys (Šulc 1934; Ruman & Hudácková 2015), the upper Miocene of France (Dell’Angelo et al. 2018b), the upper Miocene to the Pleistocene of Europe (Marquet 1984, 2002; Dell’Angelo et al. 2015a, 2018 b, 2024; this study). Remarks. Revision of recent Hanleya recorded to date in the Atlantic Ocean and Mediterranean Sea is provided by Sirenko (2014) and Sirenko et al. (2016), with two species described as new, one from the Atlantic (H. harasewychi Sirenko, 2014) and one from the Mediterranean (H. mediterranea Sirenko, 2014). These authors considered H. nagelfar (Lovén, 1846) to be a junior synonym of H. hanleyi. The morphological characters of Hanleya spp. considered in the present study are reported in Tab. 7.