Cyrtolabulus oblongus Selis, 2025, sp. nov.

Cyrtolabulus oblongus sp. nov. (Figs 26, 63N) Diagnosis. Cyrtolabulus oblongus sp. nov. belongs to the group of Malagasy Cyrtolabulus with S2 as long as T2 and propodeum without a horizontal face behind metanotum. It can be easily distinguished from all other species of this group, and in fact from most species of Cyrtolabulus, by elongate and subcylindrical mesosoma (Figs 26A, B), tegula with subtriangular posterior lobe and angled outer margin, subtriangular and wide parategula (Fig. 26C), subtriangular S1 with sides evenly diverging from base to apex, and long and translucent apical lamella of T2 with large pigmented basal digitations (Fig. 26F). The elongated habitus and general appearance could cause confusion with the genus Eumenidiopsis, represented in Madagascar by the sole species Eumenidiopsis pyriformis sp. nov., however the two genera differ profoundly in the morphology of head, clypeus, and mandibles, as already illustrated in the key to the genera. The two species further differ in numerous other characters, such as the morphology of pronotum, parategula, metanotum, propodeum, and T1. Type material examined. HOLOTYPE: ♂ labelled “CASENT / 2046420 // MADAGASCAR: Mahajanga / Prov. NP d’Ankarafantsika / Ampijoroa Sta. For. 40km 306°/ NW Andranofasika, 130m / 16°19'15"S 40°48'38"E / 26 Mar–1 Apr 2001, col. / Fisher, Griswold et al. // CASLOT 011068 / malaise in tropical dry / forest BLF3520 // Cyrtolabulus oblongus / HOLOTYPUS ♂ / Det. Marco Selis 2024 [red label]” (CAS, code CASTYPE21748). PARATYPES: Mahajanga, Foret Tsimembo 8.7 km 336°NNW Soatana, 19.021388S 44.440523E, 20 m, 21– 25.XI.2001, leg. Fisher, Griswold et al., 1♂ (CAS, code CASENT 2046325); Mahajanga, PN de Namoroka, 9.8 km 300°WNW Vilanandro, 16.466687S 45.349978E, 140 m, 4–8.XI.2002, leg. Fisher, Griswold et al., 1♂ (CAS, code CASENT 2146073); Mahajanga, PN Tsingy de Bemaraha, 2.5 km 62°ENE Bekopaka, Ankidrodroa River, 19.132232S 44.814722E, 100 m, 11–15.XI.2001, leg. Fisher, Griswold et al., 1♀ (CAS, code CASENT 2046352); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, 15.157510S 47.734166E, 97 m, 13– 19.XII.2010, leg. M. Irwin & R. Harin’Hala, 1♂ 1♀ (CAS, code CASENT 2118526; MSVI, code CASENT 2118524); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, 15.157510S 47.734166E, 97 m, 27.I– 2.II.2011, leg. M. Irwin & R. Harin’Hala, 1♀ (CAS, code CASENT 2118488); Majunga, Ambovomamy Belambo, 20 km NW of Port Berger, 15.451197S 47.613333E, 33 m, 22–27.I.2007, leg. R. Harin’Hala, M. Irwin & F. Parker, 1♂ (CAS, code CASENT 8105803); Majunga, Ambovomamy Belambo, 20 km NW of Port Berger, 15.451197S 47.613333E, 33 m, 13–15.XI.2007, leg. R. Harin’Hala, M. Irwin & F. Parker, 1♀ (CAS, code CASENT 2046252); Majunga, Ambovomamy Belambo, 20 km NW of Port Berger, 15.451197S 47.613333E, 33 m, 23.XI–8.XII.2007, leg. R. Harin’Hala, M. Irwin & F. Parker, 1♀ (CAS, code CASENT 8105934); Majunga, Analamanitra Forest, 14 km NE of Misinjo, 16.133322S 45.699989E, 2–9.X.2007, leg. M. Irwin & R. Harin’Hala, 2♀ (CAS, codes CASENT 2047013, CASENT 2047015); Majunga, Reserve forestière Beanka, 50.2 km E Maintirano, 18.026399S 44.050555E, 250 m, 19–26.X.2009, leg. B.L. Fisher et al., 1♂ (CAS, code CASENT 2166021); Tulear, Zombitse NP, near ANGAP office, 22.886509S 44.692166E, 840 m, 9–19.XI.2001, leg. R. Harin’Hala, 1♂ 1♀ (CAS, codes CASENT 2053929, CASENT 8600800); Tulear, Zombitse NP, near national road, 22.840490S 44.731155E, 825 m, 15.X–9.XI.2001, leg. R. Harin’Hala, 1♂ (MSVI, code CASENT 2153886). Description. Male holotype (Fig. 26B). Body length 5.5 mm; fore wing length 4.5 mm. Head 1.1× as wide as high. Clypeus in frontal view 1.15× as wide as long, free apical part much shorter than interocular part, apical margin shallowly emarginate and 0.25× as wide as clypeus, apical teeth wide and subtriangular with pointed apex; clypeus in lateral view weakly convex basally, then flattened to apex. Vertex 1.5× as long as ocellocular distance, very weakly convex and weakly sloping, occipital carina placed about at same level of upper limit of eye. Gena 0.65× as wide as eye at bottom of ocular sinus, somewhat bulging in upper half; occipital carina complete and sharp, shortly lamellate on gena, subparallel to outer eye margin in upper half and then smoothly bent below. F1 1.2× as long as wide and 0.95× as long as F2, F2–F3 subquadrate, F4–F8 transverse and becoming progressively wider, F9 longer than wide; F11 claw-shaped and thin, straight in dorsal view and weakly but clearly curved in lateral view, apex bluntly pointed and not reaching basal margin of F9, which is ventrally furrowed to house F11. Mesosoma 1.8× as long as wide, elongate and subcylindrical with sides converging both anteriorly and posteriorly from level of tegula. Pronotum elongate with sinuate sides, weakly concave behind humeri, which are almost right-angled and slightly pointing out, distance between pronotal carina and mesoscutum on mid-line as long as vertex; pronotal carina complete, although very fine and almost disappearing in the median third, becoming higher laterally and forming a short translucent lamella on humeri, where it is sharply bent in a right angle, then disappearing below humeri; anterior face of pronotum completely separated from dorsal faces even in the middle where the carina fades; pretegular carina present only as a very blunt trace near posterolateral corners of pronotum, replaced by a deep semicircular depression bordering pronotal lobe. Scutellum subquadrate, only slightly wider than long and with concave sides, surface evenly flattened; axillary fossa small and rounded, smaller than one ocellus. Metanotum short and sloping, divided into two faces by a robust transverse carina which is strongly bilobate, the anterior face is nearly horizontal, and the posterior face is depressed and facing the propodeal cavity. Tegula small and about as long as wide, reaching its maximum width in anterior half, subcampanulate due to abruptly curved outer margin, posterior lobe broad and somewhat acute with rounded apex, equaling parategula; parategula beak-shaped with curved apex, surface mostly flattened. Mesepisternum with a short and indistinct stretch of epicnemial carina below pleural suture, sharply separating it from epicnemium, epimeron ventrally produced in a sharp but low point. Propodeum elongate and falling just behind metanotum, in lateral view convex above and shortly produced behind metanotum and then oblique, not forming a propodeal shelf; dorsal faces of propodeum weakly and evenly convex, sloping mediolaterally, sending a short point behind metanotum but non reaching contralateral; lateral faces of propodeum flattened, clearly separated from dorsal faces but without lateral carinae; posterior face of propodeum forming a large and deep subtriangular depression with convex sides, with a longitudinal furrow originating from posterior margin of metanotum, depression clearly delimited but transitions to dorsal faces evenly rounded and not sharp; submarginal carina of propodeum forming subtriangular laterolaterally compressed lamella projections, appearing subtriangular with rounded apex in lateral view, narrower and more pointed in dorsal view. T1 elongate subconical in dorsal view, 1.4× as long as apically wide, sides markedly diverging and anterior margin subtruncate, apical margin much wider than base of tergite; T 1 in lateral view strongly and evenly convex from base to apex, except for wide preapical furrow and thickened apical margin. T2 longer than wide with subparallel sides, constricted basally and much wider apically than basally; apical margin of T2 with a long translucent orange lamella, preceded by a thickening of the tergite that sends long and blunt digitations on the basal half of the lamella. T3 lamellate similar to T2, but lamella longer and basal digitations sharper and more irregular. S1 subtriangular and covering most of the ventral surface of the segment, with only a very short basal stalk; sides markedly diverging from base to apex, separated from median shallowly depressed part by a pair of strong ridges that run subparallel to the lateral margins. S 2 in lateral view evenly convex from base to apex, less convex than T2; apical margin of S2 lamellate like on T2, but lamella apically reflexed and basal digitations sharper and more irregular. S7 with an apical broad and shallow incision laterally margined by pointed subtriangular lobes, lobes exceeding apical margin of T7 and visible also from above. Frons with deep punctures separated by 0.5–1× their diameters, interspaces flattened and shiny with sparse micropunctures, interspaces becoming wider and less micropunctate on vertex; gena with smaller and denser punctures above, becoming much sparsely punctate below; clypeus with small deep punctures separated by about their diameter, interspaces sparsely micropunctate. Mesosoma with large deep punctures and shiny interspaces; dorsal face of pronotum with very narrow interspaces; anterior face of pronotum with smaller sparse punctures; lateral faces of pronotum with punctures gradually disappearing ventrally; mesoscutum with punctures smaller than on pronotum, denser anteriorly and medially, becoming sparser posterolaterally where the interspaces are as wide as one puncture diameter and flattened; scutellum sculpted like posterior part of metanotum, but interspaces larger and posterior corners almost impunctate; metanotum with few small punctures near anterior margin; tegula shiny and sparsely micropunctate; mesepisternum sculpted similar to mesoscutum and scutellum, epimeron densely punctate; dorsal faces of propodeum with deep punctures slightly smaller than those on mesoscutum, punctures denser along margins but always clearly separated from each other and leaving some large impunctate areas on disc; lateral faces of propodeum with fine blunt striation arranged in an irregular pattern; posterior face of propo