Euspira moravica Harzhauser, Landau & Guzhov, 2025, nov. sp.

Euspira moravica nov. sp. Figs 40A–D Type material. Holotype: NHMW 1861 /0045/0153, SL: 23.0 mm, MD: 21.8 mm, Hrušovany nad Jevišovkou (Czech Republic), Figs 40D 1 –D 4. Paratypes: NHMW 1863 /0015/1011, SL: 22.5 mm, MD: 20.2 mm, Hrušovany nad Jevišovkou (Czech Republic), Figs 40A 1 –A 4. NHMW 1863 /0015/1011a, SL: 22.8 mm, MD: 21.2 mm, Hrušovany nad Jevišovkou (Czech Republic), Figs 40B 1 –B 4. NHMW 1863 /0015/1011b, SL: 23.5 mm, MD: 21.8 mm, Hrušovany nad Jevišovkou (Czech Republic), Figs 40C 1 –C 4. NHMW 1861 /0045/0153a, SL: 25.9 mm, MD: 23.9 mm, Hrušovany nad Jevišovkou (Czech Republic). NHMW 1861 /0045/0153b, SL: 25.8 mm, MD: 25.6 mm, Hrušovany nad Jevišovkou (Czech Republic). Additional material. 36 spec., NHMW 1861 /0045/1520, Hrušovany nad Jevišovkou (Czech Republic). Diagnosis. Medium sized, elongate-globose, solid shell with narrow subsutural shelf, weakly convex below, strongly convex at periphery and base; umbilicus narrow, deep, narrowed by trigonal umbilical callus fused with anterior lobe and thick parietal callus. Description. Medium sized, elongate-globose, solid shell; usually slightly higher than wide (SL/MD = ~1.1). Protoconch not preserved. Spire moderately elevated, conical; apical angle 100–112°. Suture linear, adpressed [note that the canaliculate appearance in some specimens is due to preservation]. Teleoconch whorls with very narrow, weakly convex subsutural shelf; no clear shoulder angulation. Last whorl weakly expanding, attaining ~90% of total height, weakly convex below subsutural shelf, strongly convex at periphery and base. Periphery below mid-whorl (~42%). Growth lines distinct and weakly prosocyrt in apical view, prosocline and delicate at periphery. Aperture moderately wide D-shaped, prosocline. Position of adapical tip of aperture moderately high (~82%), slightly above mid-whorl. Aperture attaining ~71–73% of total height. Columellar lip slightly concave, basal lip broad, reinforced, outer lip thin. Columellar angle ~20–31°. Opercular ridge in outer lip insignificant. Umbilicus moderately narrow (~14%), deep with prominent growth lines. Parietal callus long (~58%), narrowing umbilicus to narrow, semicircular to ovate chink. Parietal callus strongly thickened, with straight to weakly concave margin. Anterior lobe of parietal callus prominent, only slightly expanding. Funicle covered by umbilical callus. Umbilical callus fused with anterior lobe of parietal callus, forming trigonal shield-like callus. Basal fasciole weak, rounded with prominent growth lines. Color pattern not preserved. Discussion. Euspira moravica nov. sp. differs from Euspira protracta (Eichwald, 1830) in its slightly larger size, lower spire, much lower last spire whorl and narrower umbilicus with more prominent anterior lobe of the parietal callus. Euspira merlei nov. sp. has a more elongate-ovate shell and has a wider umbilicus. Similarly, Euspira vinitor nov. sp. is larger, has a higher spire, and a larger umbilicus. Paleoenvironment. The accompanying fauna suggests shallow marine environments (own data, M.H.). Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Hrušovany nad Jevišovkou (hoc opus).