Nyctimus quadripunctatus Dhiya'ulhaq & Benjamin 2025, sp. nov.

Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin sp. nov. Figs 9, 10, 11, 12 Etymology. The specific epithet is taken from Latin, referring to the four white spots on the opisthosoma. Type material. Holotype. Indonesia – Sumatra, Jambi Province • ♂; Batang Hari, Hutan Harapan Conservation Area; 02°11'15.3"S, 103°20'36.0"E; elev. 69 m; 7 Aug. 2013; J. Drescher leg.; canopy fogging in rainforest; GOET 2013_HF 4.2_AraThom 001 N_001 (to be transferred to MZB). Paratypes. Indonesia – Sumatra, Jambi Province • 1 ♂ 1 ♀; Batang Hari, Hutan Harapan Conservation Area; 02°10'42.4"S, 103°19'58.2"E; elev. 54 m; 21 Jul. 2013; J. Drescher leg.; canopy fogging in rainforest; GOET 2013_HF 3.2_AraThom 001 N_001–2 (to be transferred to MZB) • Malaysia – Sabah State • 1 ♂; Kinabalu National Park, Poring Hot Springs; 05°59'N, 116°42'E; elev. 600–700 m; 28 Mar. 1998; A. Floren leg.; canopy fogging on Barringtonia tree, in primary forest; RMNH RMNH.ARA.17858. • 1 ♂; Kota Marudu, Kampong Sorinsim; 06°06'N, 116°50'E; elev. 600–700 m; 2 Mar. 1997; A. Floren leg.; canopy fogging in 15 years old secondary forest; RMNH RMNH.ARA.17860. Other material examined. Malaysia – Sabah State • 1 ♂; Kinabalu National Park, Poring Hot Springs; 05°59'N, 116°42'E; elev. 600–700 m; 27 Feb. 1996; A. Floren leg.; canopy fogging on Aporosa subcaudata tree, in primary forest; RMNH RMNH.ARA.17857. Diagnosis. Males are easily distinguished from other congeners by the anvil-shaped RTA ending in two acute lobes (Fig. 11 A, B). Females can be distinguished from N. bistriatus by the much thinner, shorter CD and paired CO (Figs 10 C, D, 11 C, D; vs single CO in N. bistriatus, Figs 2 C, 3 C), and from N. rendang, sp. nov., by the spermatheca lacking a prominent posterior extension (Figs 10 D, 11 D; vs Figs 14 D, 15 D). Description. Male (holotype 2013 _ HF 4.2 _ AraThom 001 N _ 001; Figs 9, 11 A, B). Total length 2.95. Prosoma length 1.35; width 1.23. Opisthosoma length 1.60; width 1.22. Diameter of eyes: AME 0.07; ALE 0.16; PLE 0.16; PME 0.06. Interdistances between eyes: AME – AME 0.38; AME – ALE 0.08; ALE – ALE 0.68; PME – PME 0.34; PME – PLE 0.32; ALE – PLE 0.19; AME – PME 0.18; PLE – PLE 0.98. Clypeus height 0.00. Leg measurements: leg I 4.25 (1.19, 0.49, 1.06, 0.90, 0.61); leg II 3.89 (1.11, 0.42, 0.90, 0.81, 0.65); leg III 2.63 (0.86, 0.35, 0.57, 0.49, 0.36); leg IV 2.74 (0.83, 0.27, 0.66, 0.52, 0.46). Prosoma cuboid, almost as long as wide, box-shaped, weakly granulated, highest at mid-point, light brown; lateral margins furnished with a row of long macrosetae. Lateral eyes large, situated on tubercles on the lateral edge of the prosoma. Chelicerae, endites, and labium light brown. Sternum light brown. Legs I light brown; others yellow-brown, with sparse spines and black setae. Opisthosoma oval, brown, with two pairs of large white spots. Spinnerets brown. Palp (Figs 9 C, D, 11 A, B): cymbium oval. Embolus sickle-shaped; base slightly wrinkled. Tegular hood very wide. RTA large, rectangular, anvil-shaped; dorsally bifurcated, ending into two long, acute lobes, hidden behind a brush of setae; VTA hook-shaped attached medially on RTA. Female (paratype 2013 _ HF 3.2 _ AraThom 001 N _ 002; Figs 10, 11 C, D). Total length 4.81. Prosoma length 1.95; width 1.77. Opisthosoma length 2.86; width 2.65. Diameter of eyes: AME 0.09; ALE 0.20; PLE 0.18; PME 0.07. Interdistances between eyes: AME – AME 0.53; AME – ALE 0.14; ALE – ALE 0.97; PME – PME 0.52; PME – PLE 0.39; ALE – PLE 0.23; AME – PME 0.24; PLE – PLE 1.37. Leg measurements: leg I 5.47 (1.66, 0.57, 1.34, 1.13, 0.77); leg II -; leg III 3.69 (1.22, 0.47, 0.87, 0.63, 0.50); leg IV 4.12 (1.36, 0.46, 0.96, 0.76, 0.58). Habitus as in males, except prosoma and leg I reddish brown. Epigynum (Figs 10 C, D, 11 C, D): exterior plate large, hexagonal-shaped with wide anterior side. Atrium conspicuous, inverted heart-shaped with a separation in the middle. CO diagonally oriented, posteriorly facing. CD arch-shaped. Spermatheca elongated, peanut-shaped, with a small bump anteriorly. Distribution. Malaysia (Borneo: Sabah); Indonesia (Sumatra: Jambi) (Fig. 19). Remarks. Specimens from Borneo show slight differences in shape of embolus and dorsal tip of RTA (Fig. 12 B, C).