Stethorrhagus naja Xavier, Brescovit & Bonaldo, 2025, sp. nov.

Stethorrhagus naja sp. nov. urn:lsid:zoobank.org:act: 5DC82360-F963-408B-B00B-E0673F637CE2 Figs 40–42, 50 Diagnosis Males of S. naja sp. nov. resemble those of S. chalybeius by the bifid VL, with dVL protuding and thick modified hairs only present on the vVL, dVL glabrous (Fig. 41B–C), differing by the large finger-shaped SePP as long as the embolus, with a blunt tip (Fig. 41A) (SePP smaller than the embolus, comma-shaped, tip pointed in S. chalybeius – Bonaldo & Brescovit 1994: fig. 16a). Females resemble those of S. bradypus sp. nov. by the epigynum with VEP, CO disposed anteriorly in relation to SI, with straight posterior margin (Fig. 41D), differing by the VEP placed posteriorly on the epigynal plate; SI not extended anteriorly, below the level of CO (Figs 41D–E, 42E–F) (VEP placed medially on epigynal plate; SI extended anteriorly, with long FDs inserted on the SI’s extension, (beyond the level of the CO in S. bradypus – Figs 38D–E, 39D–E). Etymology The specific name is a noun in apposition in reference to the elapid genus Naja Laurenti, 1768, since the anterior sector of the epigynum (as in Fig. 41D) resembles a cobra with spread hood. Material examined Holotype COLOMBIA • ♂; Cundinamarca, Parque Nacional Natural de Chingaza; [4°0′ N, 73°30′ W]; elev. 3150 m; 16 Dec. 1997; E. Niao leg.; pitfall; ICN-Ar 6477. Paratypes COLOMBIA – Cundinamarca • 1 ♀; same data as for holotype; ICN-Ar 13743 • 1 ♂; same data as for holotype; 24 Jan. 1998; Guarda Bosques leg.; ICN-Ar 6470. Description Male (holotype – ICN-Ar 6477) COLORATION. Carapace, chelicerae, endites and sternum red-brown (Fig. 40A). Labium orange. Legs brown. Abdomen gray with dorsal muscular impression orange (Fig. 40A). Sternum with deep sternal excavations (Fig. 40C). MEASUREMENTS. Total length 5.40. Carapace 2.47 long, 1.85 wide. Clypeus 0.22. Leg measurements: I: femur 1.93/ patella 0.77/ tibia 1.53/ metatarsus 1.49/ tarsus 1.19/ total 6.91; II: 1.80/ 0.76/ 1.35/ 1.42/ 1.09/ 6.42; III: 1.55/ 0.64/ 1.08/ 1.40/ 0.82/ 5.49; IV: 2.07/ 0.80/ 1.65/ 2.09/ 0.93/ 7.54. Eye diameters: AME 0.13, ALE 0.11, PME 0.12, PLE 0.11. Chelicerae 1.15 long, with three promarginal teeth and five retromarginal denticles. LEG SPINATION. I – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. II – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. III – femur d1-1-0, p0, r0; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p0-1-0, r0-1-0, v2-2-1. IV – femur d1-1-0, p0, r0-0-1; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p1-1-0, r1-1-0, v2-2-1. PALP. Retrolateral surface of femur with a cluster of modified hairs, vVL with modified hairs, dVL glabrous, well-developed, subrectangular, vVL larger than dVL, AS absent, DL absent, VTA slightly broader than long in ventral view, spermophor situated prolaterally, TP1 small, triangular, TP2 absent, SePP well-developed, finger-shaped, tip blunt, pointed retrolaterally, embolus long, not curved, prongs short, not curved (Figs 41A–C, 42A–C). Female (paratype – ICN-Ar 13743) COLORATION. As in male, except abdomen totally dark gray (Fig. 40B). Sternum with sternal excavations as in male (Fig. 40D). MEASUREMENTS. Total length 6.97. Carapace 2.70 long, 2.00 wide. Clypeus 0.22. Leg measurements: I: femur 1.99/ patella 0.94/ tibia 1.50/ metatarsus 1.49/ tarsus 1.08/ total 7.00; II: 1.92/ 0.90/ 1.34/ 1.45/ 1.14/ 6.75; III: 1.65/ 0.79/ 1.15/ 1.36/ 0.88/ 5.83; IV: 2.21/ 0.91/ 1.86/ 2.24/ 0.96/ 8.18. Eye diameters: AME 0.13, ALE 0.11, PME 0.11, PLE 0.12. Chelicerae 1.34 long, with three promarginal teeth and five retromarginal denticles. LEG SPINATION. I – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. II – femur d1-1-0, p0, r0; tibia d0, p0, r0, v0-1p-0; metatarsus d0, p0, r0, v2-2-0. III – femur d1-1-0, p0, r0; tibia d0, p0-1-0, r0-1-0, v1p-2-2; metatarsus d0, p1-1-0, r1-1-0, v2-2-1. IV – femur d1-1-0, p0, r0; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p1-1-1, r1-1-1, v2-2-1. EPIGYNUM. CO disposed anteriorly in relation to SI, CO with straight posterior margin, VEP placed posteriorlly on epigynal plate, PVP wider than long, covering entire SI, PVP as wide as four times the distance between anterior and posterior margins, PVP with slightly procurved notch along anterior margin, copulatory duct as long as two times the distance between anterior and posterior margins of PVP (Figs 41D–E, 42D–E). Distribution Known only from the type locality (Fig. 50).

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Zenodo (CERN European Organization for Nuclear Research)
Published
2025-09-15
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https://doi.org/10.5281/zenodo.17226762
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Geochemistry and Geologic Mapping
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article

Stethorrhagus naja Xavier, Brescovit & Bonaldo, 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Geochemistry and Geologic Mapping
article

Stethorrhagus naja Xavier, Brescovit & Bonaldo, 2025, sp. nov.

article en

Abstract

Stethorrhagus naja sp. nov. urn:lsid:zoobank.org:act: 5DC82360-F963-408B-B00B-E0673F637CE2 Figs 40–42, 50 Diagnosis Males of S. naja sp. nov. resemble those of S. chalybeius by the bifid VL, with dVL protuding and thick modified hairs only present on the vVL, dVL glabrous (Fig. 41B–C), differing by the large finger-shaped SePP as long as the embolus, with a blunt tip (Fig. 41A) (SePP smaller than the embolus, comma-shaped, tip pointed in S. chalybeius – Bonaldo & Brescovit 1994: fig. 16a). Females resemble those of S. bradypus sp. nov. by the epigynum with VEP, CO disposed anteriorly in relation to SI, with straight posterior margin (Fig. 41D), differing by the VEP placed posteriorly on the epigynal plate; SI not extended anteriorly, below the level of CO (Figs 41D–E, 42E–F) (VEP placed medially on epigynal plate; SI extended anteriorly, with long FDs inserted on the SI’s extension, (beyond the level of the CO in S. bradypus – Figs 38D–E, 39D–E). Etymology The specific name is a noun in apposition in reference to the elapid genus Naja Laurenti, 1768, since the anterior sector of the epigynum (as in Fig. 41D) resembles a cobra with spread hood. Material examined Holotype COLOMBIA • ♂; Cundinamarca, Parque Nacional Natural de Chingaza; [4°0′ N, 73°30′ W]; elev. 3150 m; 16 Dec. 1997; E. Niao leg.; pitfall; ICN-Ar 6477. Paratypes COLOMBIA – Cundinamarca • 1 ♀; same data as for holotype; ICN-Ar 13743 • 1 ♂; same data as for holotype; 24 Jan. 1998; Guarda Bosques leg.; ICN-Ar 6470. Description Male (holotype – ICN-Ar 6477) COLORATION. Carapace, chelicerae, endites and sternum red-brown (Fig. 40A). Labium orange. Legs brown. Abdomen gray with dorsal muscular impression orange (Fig. 40A). Sternum with deep sternal excavations (Fig. 40C). MEASUREMENTS. Total length 5.40. Carapace 2.47 long, 1.85 wide. Clypeus 0.22. Leg measurements: I: femur 1.93/ patella 0.77/ tibia 1.53/ metatarsus 1.49/ tarsus 1.19/ total 6.91; II: 1.80/ 0.76/ 1.35/ 1.42/ 1.09/ 6.42; III: 1.55/ 0.64/ 1.08/ 1.40/ 0.82/ 5.49; IV: 2.07/ 0.80/ 1.65/ 2.09/ 0.93/ 7.54. Eye diameters: AME 0.13, ALE 0.11, PME 0.12, PLE 0.11. Chelicerae 1.15 long, with three promarginal teeth and five retromarginal denticles. LEG SPINATION. I – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. II – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. III – femur d1-1-0, p0, r0; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p0-1-0, r0-1-0, v2-2-1. IV – femur d1-1-0, p0, r0-0-1; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p1-1-0, r1-1-0, v2-2-1. PALP. Retrolateral surface of femur with a cluster of modified hairs, vVL with modified hairs, dVL glabrous, well-developed, subrectangular, vVL larger than dVL, AS absent, DL absent, VTA slightly broader than long in ventral view, spermophor situated prolaterally, TP1 small, triangular, TP2 absent, SePP well-developed, finger-shaped, tip blunt, pointed retrolaterally, embolus long, not curved, prongs short, not curved (Figs 41A–C, 42A–C). Female (paratype – ICN-Ar 13743) COLORATION. As in male, except abdomen totally dark gray (Fig. 40B). Sternum with sternal excavations as in male (Fig. 40D). MEASUREMENTS. Total length 6.97. Carapace 2.70 long, 2.00 wide. Clypeus 0.22. Leg measurements: I: femur 1.99/ patella 0.94/ tibia 1.50/ metatarsus 1.49/ tarsus 1.08/ total 7.00; II: 1.92/ 0.90/ 1.34/ 1.45/ 1.14/ 6.75; III: 1.65/ 0.79/ 1.15/ 1.36/ 0.88/ 5.83; IV: 2.21/ 0.91/ 1.86/ 2.24/ 0.96/ 8.18. Eye diameters: AME 0.13, ALE 0.11, PME 0.11, PLE 0.12. Chelicerae 1.34 long, with three promarginal teeth and five retromarginal denticles. LEG SPINATION. I – femur d1-1-0, p0-0-1, r0; tibia d0, p0, r0, v0-2-0; metatarsus d0, p0, r0, v2-2-0. II – femur d1-1-0, p0, r0; tibia d0, p0, r0, v0-1p-0; metatarsus d0, p0, r0, v2-2-0. III – femur d1-1-0, p0, r0; tibia d0, p0-1-0, r0-1-0, v1p-2-2; metatarsus d0, p1-1-0, r1-1-0, v2-2-1. IV – femur d1-1-0, p0, r0; tibia d0, p1-1-0, r1-1-0, v1p-1p-2; metatarsus d0, p1-1-1, r1-1-1, v2-2-1. EPIGYNUM. CO disposed anteriorly in relation to SI, CO with straight posterior margin, VEP placed posteriorlly on epigynal plate, PVP wider than long, covering entire SI, PVP as wide as four times the distance between anterior and posterior margins, PVP with slightly procurved notch along anterior margin, copulatory duct as long as two times the distance between anterior and posterior margins of PVP (Figs 41D–E, 42D–E). Distribution Known only from the type locality (Fig. 50).

Zenodo (CERN European Organization for Nuclear Research)
Museu Paraense Emílio Goeldi (BR), Instituto Butantan (BR)
Openalex Percentile: Top 26%
Geochemistry and Geologic Mapping
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