Chimalfus discoideus Larsen & Solodovnikov & Reyes-Hernández 2025, comb. nov

3.1.6. Chimalfus discoideus (Fairmaire and Germain, 1862), comb. nov Figs. 1C, 2C and 5 and 6. Fairmaire and Germain (1862): 427 (as H. discoideus, original description); Reyes-Hern´andez et al. (2024): 38 (as Sphingoquedius discoideus, taxonomic notes). Type material examined: Lectotype: male, pinned, with labels as follows: Santiago quillota Concepcion //Coll. et det A. Fauvel Quedius discoideus Fairmaire and Germ. R.I.Sc.N.B. 17.479 //Lectotype [red label]// Loncovilius (Lienturius) discoideus 1964 det. Coiff. et Saiz [IRSNB]; paratypes: Santiago quillota Concepcion // discoideus Fairm. Germ. // R.I.Sc.N.B. 17.479 Coll. et det A. Fauvel//paralectotype [red label]// Loncovilius (Lienturius) discoideus 1964 det. Coiff. et Saiz [1♀ IRSNB]. Other material examined: see Supplementary File 1. 3.1.7. Redescription Measurements ♀ [min–max (average); n = 5]: FBL = 3.09–3.59 (3.30); TL = 5.96–6.90 (6.39); HW = 0.68–0.81 (0.65); HL = 0.66–0.90 (0.67); HW/HL = 0.84–1.23 (0.98); PW = 1.04–1.18 (0.94); PL = 0.95–1.20 (0.86); PW/PL = 0.86–1.21 (1.11); EW = 1.45–1.49 (1.22); EL = 1.36–1.54 (1.22); PW/HW = 1.30–1.74 (1.47). Measurements Ƌ [min–max (average); n = 5]: FBL = 3.11–3.66 (3.29); TL = 6.22–7.15 (6.67); HW = 0.76–0.80 (0.79); HL = 0.76–0.83 (0.80); HW/HL = 0.94–1.05 (0.99); PW = 1.14–1.26 (1.19); PL = 0.99–1.09 (1.03); PW/PL = 1.12–1.18 (1.15); EW = 1.36–1.87 (1.53); EL = 1.34–1.75 (1.46); PW/HW = 1.42–1.63 (1.51). Body including legs dark brown to almost black; apical margins of head and pronotum yellowish; occasionally legs yellowish-brown. Head as wide as long; dorsally and ventrally glossy with few micropunctures; frons with microsculpture of concentric transverse waves transforming into meshed or isodiametric towards the apex. Nuchal ridge fused with infraorbital ridge; eyes large (EYL/HL = x = 0.63); distance between eyes about 0.9 times the length of eyes in both sexes. Antennal proportions as follows: a1 at least 1.65 times longer than a2; a2 and a3 subequal in length; a4 to a10 subequal in length; a11 more than 1.6 times longer than a10, in both sexes. All antenomeres longer than wide. Pronotum slightly wider than long, convex, evenly curved; flexible postcoxal hypomeral extension rounded, as wide as long, uninterrupted by inferior line. Elytra distinctly longer than pronotum. Mesosternum with eleven to twelve macrosetae. Metathoracic wings well developed. Protarsomeres 1–3 distinctly wider than long. Metatarsi with first tarsomere about as long as apical tarsomere; all apical tarsomeres with empodial setae longer or subequal to tarsal claws; metacoxae with four or more spines on posterior margin; metatrochanter rounded at apex. Abdominal tergite II with distinct row of setiferous punctures along apical margin; tergite VII with white fringe along posterior edge. Male sternite VIII with broad and shallow U-shaped emargination. Aedeagus (Fig. 5D and E), its total length ~ 1.2 mm. Median lobe tube shaped; paramere extends over apex of median lobe, narrower at midpoint but wider towards apex; paramere in lateral view almost spoon-like in shape. Comparison. This species is differentiated from other congeners by the following combination of characters: length of the eyes more than half the length of the head (EYL/HL = x = 0.63), frons apically with isodiametric mesh microsculpture, metathoracic wings well developed and tergite VII with white fringe along posterior edge. This species is significantly distinguished from others by the shape of the head and pronotum chaetotaxy. In morphospace, it shows wide interspecific variation, overlapping with C. brunneus in pronotum shape under allometric effects. When corrected for size, the morphospace shows overlaps with the head and pronotum of L. nana, which share similar setae configurations but are differentiated by size (Fig. 9; see section 3.2 for more details). Distribution. C. discoideus is distributed across central and southern Chile, from the Coquimbo Region to the Los Lagos Region. It inhabits both the Chilean Matorral and Valdivian Temperate Forest ecoregions.