Eutyphoeus Michaelsen 1900

Updated key to the species of Indian Eutyphoeus Michaelsen, 1900 (revised after Julka 1988 and Tiwari et al. 2021) 1. Holandric; seminal vesicles in ix and xii .......................................................................................... 2 – Metandric; seminal vesicles in xii .................................................................................................. 16 2. Spermathecal pores at a or ab ........................................................................................................... 3 – Spermathecal pores at b or lateral to b .............................................................................................. 5 – Spermathecal pores at bc .................................................................................................................. 8 3. Spermathecae unidiverticulate ............................ E. manipurensis manipurensis Stephenson, 1921 – Spermathecae bidiverticulate ............................................................................................................ 4 – Spermathecae polydiverticulate ............................................. E. quadripapillatus Michaelsen, 1907 4. Spermathecal pore at a ............................ E. mizoramensis Julka, Ramanujam & Lalthanzara, 2005 – Spermathecal pore at ab .................................................................................................................... 9 5. Vestibulate ......................................................................................................................................... 6 – Avestibulate ......................................................................... E. morehensis Tiwari & Yadav sp. nov. 6. Lateral intestinal caecae present ....................................................................................................... 7 – Lateral intestinal caecae absent ....................... E. tawi Tiwari, Lone, Thakur, James & Yadav, 2021 7. Genital markings present ................................................................................................................ 13 – Genital markings absent .......................................................................... E. marmoreus Gates, 1933 8. Spermathecae bidiverticulate .......................................................... E. annandalei Michaelsen, 1907 – Spermathecae polydiverticulate ..................................................... E. incommodus (Beddard, 1901) 9. Clitellum annular ............................................................................................................................ 10 – Clitellum not annular .......................................................................................................................11 10. Genital marking gland absent, lateral intestinal caeca dorsally directed ............................................ ....................................................... E. phawngpuiensis Tiwari, Lone, Thakur, James & Yadav, 2021 – Genital marking gland present, lateral intestinal caeca ventrally directed ......................................... ........................................................................................... E. lamdengensis Tiwari & Yadav sp. nov. 11. Penial setae present ........................................................................................................................ 12 – Penial setae absent ................................................................ E. shiroensis Tiwari & Yadav sp. nov. 12. Male pore under cuneate wedge-shaped papillae with fimbriated margin at b ................................... ........................................................................................... E. kashongensis Tiwari & Yadav sp. nov. – Male pore on circular porophores centered at b ................. E. ukhrulensis Tiwari & Yadav sp. nov. 13. Genital marking gland around spermathecal duct in vii present .................................................... 14 – Genital marking gland around spermathecal duct in vii absent ...................................................... 15 14. Vestibula pentagonal shape, penial setae with blunt tip ...................................................................... ..................................................................................... E. yangoupokpensis Tiwari & Yadav sp. nov. – Vestibula not-pentagonal shape, penial setae with pointed tip............................................................. ...................................................................................... E. tengnoupalensis Tiwari & Yadav sp. nov. 15. Preclitellar genital marking present ..................................... E. lokchaensis Tiwari & Yadav sp. nov. – Preclitellar genital marking absent ..................................... E. baronongini Tiwari & Yadav sp. nov. 16. Avestibulate .................................................................................................................................... 17 – Vestibulate ....................................................................................................................................... 30 17. Spermathecae bidiverticulate .......................................................................................................... 18 – Spermathecae polydiverticulate ...................................................................................................... 28 18. Lateral intestinal caecae present ..................................................................................................... 19 – Lateral intestinal caecae absent ...................................................................................................... 24 19. Lateral intestinal caecae dorsally directed ...................................................................................... 20 – Lateral intestinal caecae ventrally directed ..................................................................................... 21 20. Genital marking glands present ........................................................................ E. lippus Gates, 1934 – Genital marking glands absent............. E. rihnimensis Rodingpuia, Lalthanzara, Julka, Chawngthu, Mathipi, Lalfelpuii, Kumar, Gurusubramaniun, Lalchhandama, Vanlalrovi & Chhakchhuak, 2024 21. Female pore single, on left side ...................................................................................................... 22 – Female pore paired ......................................................................................................................... 23 22. Penial setae ornamented, with tapering tip ............................................... E. gigas Stephenson, 1917 – Penial setae without ornamentation, with spoon shaped tip ............................................................... .......................................................................... E. serei Tiwari, Lone, Thakur, James & Yadav, 2021 23. Penial setae shaft slightly curved, tip thickened, pointed and claw-shaped ....................................... ............................................................................................................ E. turaensis Stephenson, 1920 – Penial setae serpentine, tip spoon-shaped with a small bulbous end .................................................. ................................................................................................ E. kwathensis Tiwari & Yadav sp. nov 24. Spermathecal pores at ab ................................................................................................................ 25 – Spermathecal pores lateral to b ....................................................................................................... 26 25. Genital markings preclitellar, segmental ...................................... E. comillahnus Michaelsen, 1907 – Genital markings postclitellar, intersegmental .............................................. E. festivus Gates, 1938 26. Genital markings paired, segmental on xvi .................................................................................... 27 – Genital markings unpaired, intersegmental on xv/xvi ....................... E. scutarius Michaelsen, 1907 27. Sessile genital marking glands present ............................................................................................... ............................................................. E. dhubriensis Ahmed, Julka, Banerjee & Marimuthu, 2023 – Sessile genital marking glands absent .......................................... E. assamensis Stephenson, 1926 * 28. Seminal vesicles in xii very long, extending posteriorly into xxxiii .................................................. ................................................................................................... E. pharpingianus Michaelsen, 1907 – Seminal vesicles in xii shorter, at the most extending into xx ........................................................ 29 29. Genital marking intraclitellar, on xvi ................................................ E. nainianus Michaelsen, 1907 – Genital marking intra and post-clitellar, on xiv–xv, xviii/xix–xxi/xxii ............................................... .......................................................................................................... E. nepalensis Michaelsen, 1907 30. Univestibulate ................................................................................................................................. 31 – Bivestibulate ................................................................................................................................... 32 31. Spermathecal pores at ab; penes conical .......................................................... E. kherai Julka, 1978 – Spermathecal pores at median half of bc; penes annular ...................... E. gammiei (Beddard, 1888) 32. Penes annular .......................