Sphaeronectria Darmostuk, Etayo & Flakus 2025, gen. nov.

Sphaeronectria Darmostuk, Etayo & Flakus, gen. nov. MycoBank MB 858389. Etymology: Referring to the globose ascomata. Type species: Sphaeronectria lichenophila (Speg.) Darmostuk, Etayo & Flakus Ascomata perithecioid, superficial, globose to subglobose, constricted laterally or apically, yellowish to orange, with tiny tomentum on the lower part of ascomata, 150–250 μm diam., covered by abundant septate hyaline hyphae, 50–100 µm long. Ascomatal wall formed by several layers of cells, hyaline to pale yellowish in the outer part, up to 20 μm thick, superficially paraplectenchymatous, KOH–. Asci unitunicate, subcylindrical, 4-spored, without apical thickness. Ascospores hyaline, 1-septate, ellipsoid, with pointed ends, slightly constricted at the septa, smooth-walled. Conidiomata not observed. Notes: Previously the type species of Sphaeronectria (S. lichenophila) was included in the genus Nectriopsis, typified by N. violacea, but based on our phylogenetic results, S. lichenophila showed a different phylogenetic position in Paranectriaceae, whereas Nectriopsis is a member of Bionectriaceae. Currently, the genus is monotypic.