Heinrichiellus natgeo Ranjith & Broad 2025, sp. nov.

Heinrichiellus natgeo Ranjith & Broad sp. nov. Material examined. Holotype: • female, India: Arunachal Pradesh, Yingku, 5.v.2022, Yellow pan trap, coll. Ranjith A. P. (ZSIK: ZSI/WGRC/I.R.-INV.28943). Paratype • 1 male with same data as holotype (ZSIK: ZSI/WGRC/I.R.-INV.28944) (with following DNA data: sample ID CCDB-44298-H 01, BOLD Process ID BBTH 5022-22; GenBank accessions: COI PQ 858774, 16 S PQ 873015, 28 S PQ 873014). Holotype. ♀, body length 10.7 mm, fore wing length 7.7 mm. Description of female. Head. Head 1.2 × as wide as high in anterior view, 1.5 × as wide as long in dorsal view, minutely punctate (Fig. 5 B, C); ocellar-ocular length (OOL) / postero-ocellar length (POL) = 1.0. Frons slightly convex above each antennal socket (Fig. 5 C); frons granulate with small and distinct tubercle between antennal sockets (Fig. 5 C); antennal sockets large and protruding anteriorly in dorsal view (Fig. 5 C); face 1.4 × as wide as high, smooth, irregularly sculptured below antennal socket (Fig. 5 B); inner margins of compound eyes slightly diverging ventrally (Fig. 5 B); clypeus strongly convex and transverse, 1.6 × as wide as high, with medially concave apical margin (Fig. 5 B); anterior tentorial pits large (Fig. 5 B); labrum very wide, 0.6 × as wide as clypeus, with straight apical margin, protruding from under clypeus (Fig. 5 B). Mandible slender, 2.8 × as long as wide at base, gradually narrowed from base to apex, with indistinct lower tooth (Fig. 5 B, D). Malar space granulate, 3.2 × as wide as base of mandible (Fig. 5 D). Occipital carina complete, joining hypostomal carina above base of mandible (Fig. 5 D); dorso-median part of occipital carina evenly arched (Fig. 5 C). Maxillary palp long, reaching epicnemial carina (Fig. 5 D); fifth segment 5.0 × as long as mandible base. Dorsal length of eye 1.7 × as long as temple. Antenna with 36 antennomeres. Flagellum bristle-shaped, very long and slender (Fig. 5 A); apical flagellomere conical; first flagellomere 6.2 × as long as apical width and 1.8 × as long as second one. Mesosoma. Mesosoma 1.6 × as long as high. Collar of pronotum long (Fig. 5 D – F). Pronotum strigose on coriaceous surface laterally (Fig. 5 D, E). Mesoscutum strongly convex (Fig. 5 E), with notaulus reaching middle of mesoscutum (Fig. 5 F); surface of mesoscutum granulate with scattered punctures except longitudinally strigose postero-median portion (Fig. 5 F). Epicnemial carina complete (Fig. 5 E); subtegular ridge sharp (Fig. 5 E); impression below speculum weak (Fig. 5 E); sternaulus distinct, crenulate, sharp, almost reaching base of mid coxa (Fig. 5 E); posterior transverse carina of mesosternum absent; surface of mesopleuron obliquely strigose, smooth anteriorly, speculum slightly polished (Fig. 5 E). Scutellum highly elevated above metanotum (Fig. 5 E), with crenulate lateral carina reaching apex and dorsal surface almost flat (Fig. 5 F); hind margin of metanotum with a triangular projection on each side of metanotum (Figs 5 F, 6 A). Juxtacoxal carina distinct, complete (Fig. 5 E). Propodeum in profile with triangular area basalis flat and evenly slanted from base of area superomedia (Fig. 5 E); regular carinae of propodeum complete (Fig. 6 A); area superomedia hexagonal, 1.4 × as long as wide (Fig. 6 A); apex of area dentipara with very strongly developed, long and sharp apophysis, almost perpendicular to surface of propodeum, apophysis almost straight (Fig. 6 A); surface of propodeum smooth (Fig. 6 A); area petiolaris indistinct laterally, transversely strigose (Fig. 6 A). Propodeal spiracle 1.4 × as long as wide. Legs. Legs very slender and long (Fig. 5 A); all legs longer than fore wing. Hind femur and tibia 4.5, 7.0 × as long as maximum width respectively; ratio of length of tibiae fore: mid: hind = 1.0: 1.1: 1.6. Wings. Fore wing 5.9 mm long; areolet 1.5 × as wide as long, 1.5 × as high as 3 rs-m, 0.9 × as high as 2 m-cu, pentagonal and sides nearly parallel, slightly narrowing anteriorly (Fig. 6 E); 1 cu-a opposite M & RS (Fig. 6 E); ramulus absent (Fig. 6 E). Hind wing with cu-a one-fourth as long as first abscissa of CU. Metasoma. Metasoma mostly coriaceous (Fig. 6 B – D). First tergite in profile almost straight, slightly bent downward at base of postpetiole (Fig. 6 B), smooth anteriorly, irregularly sculptured between latero-median carinae; in dorsal view 5.0 × as long as wide at base (Fig. 6 C); petiole very long, rounded in cross-section, of approximately equal width and height (Fig. 6 B, C); in dorsal view petiole slightly broadened posteriorly; latero-median carina present only on anterior portion of postpetiole (Fig. 6 C). Second tergite in dorsal view 2.6 × as long as wide anteriorly, distinctly punctate (Fig. 6 D). Gastrocoelus slightly impressed, in form of longitudinal groove (Fig. 6 D). Thyridium oval, distant from base of second tergite by 0.3 × its length (Fig. 6 D). Tergites 3–6 smooth with laterotergites separated by distinct creases (Fig. 6 B). Sternites 2–5 laterally sclerotized and middle part not sclerotized (Fig. 6 B). Hypopygium with apical margin rounded and longitudinally folded in middle. Ovipositor sheath with apex pointed, protruding beyond apex of metasoma (Fig. 6 B). Colour. Body black except following: clypeus apically, maxillary and labial palps, face antero-laterally, scape and pedicel, flagellomeres 7–11, frons laterally, pronotum antero-laterally, mesopleuron anteriorly, scutellum posterior half, propodeum, metapleuron, fore and mid legs except tarsi, hind coxa, femur, tibia proximal half, basitarsus distal half, tarsomeres 2–4, first metasomal tergite posterior 1 / 3, second metasomal tergite posterior half, metasomal tergites 6–7, ovipositor yellow. Male. Similar to female except: body length 5.1 mm, white inner orbits and clypeus laterally, apically; dorsal inner orbit patch narrowly separated from wider white line below antennal sockets. T 1 and hind coxa (rest of hind legs missing) largely dark brown. Gonostyle narrow, ‘ finger-like’. Distribution. Oriental region (India). Etymology. We dedicate the new species to the National Geographic Society (NGS) for their outstanding support for the Siang Expedition in which specimens of the new species were collected. It’s a humble dedication to the society in recognition of their committed support to explore, illuminate and protect the wonders of our world through research. The epithet natgeo is treated as a noun in apposition. Comparative diagnosis. The new species is distinctive and unique as the fore wing areolet is very large, hardly narrowing anteriorly. The propodeal spiracle is also shorter than in other Heinrichiellus, 1.4 × as long as wide. Superficially this species resembles Heinrichiellus brevispinus sp. nov. but can additionally be separated by the length of the propodeal apophyses and the shape of the area superomedia.