Subprotelater miriensis Arimoto & Kaliang & Mizuno & Meleng & Kishimoto-Yamada & Itioka 2025, sp. nov.

Subprotelater miriensis Arimoto, sp. nov. (Figures 4–6) Type material. Malaysia, Sarawak, Miri, Lambir Hills National Park, by ultraviolet light traps. Holotype. Male (AA2739), 17 m, 23–24 VIII 1998. Paratypes. 1 male (AA2738), 17 m, 14–15 XII 1996; 1 male (AA2740), 35 m, 18–19 XII 1998; 1 male (AA2741), 17 m, 17–18 V 1999; 1 male (AA2742), 35 m, 15–16 II 2002. Male. Diagnosis. This species is characterized by the following features: punctures large and umbilicate; interspaces between punctures generally shorter than puncture diameter; body brown; antennae and legs orange; elytron with four sub-circular yellow spots; setae yellow; pronotum 1.3 × longer than wide; prosternal process 2.0– 2.3 × longer than procoxal cavity length in ventral view; mesal posterior angle of hypomeron obtuse, not protruding; posterior part of scutellar shield visible in lateral view; elytron 5.2–5.5 × longer than wide, 2.0–2.2 × longer than pronotal length; metepisternum narrower than width of elytral epipleuron; basal struts 0.3 × total length of median lobe; apices of parameres tapering to apex. Measurements (5 specimens; holotype in parentheses). BL: 4.61–6.13 (4.61), BW: 1.11–1.54 (1.11), MAE: 0.90–1.11 (0.90), MBE: 0.54–0.66 (0.54), OI: 160–171 (169), PL: 1.41–1.99 (1.41), PML: 1.28–1.75 (1.28), PW: 1.10–1.56 (1.10), PAW: 0.87–1.12 (0.87), PLI: 128–129 (129), PWI: 126–146 (126), EL: 3.01–4.02 (3.01), EW: 0.55–0.75 (0.55), EI: 521–545 (545), BI: 197–216 (213). Description. Body (Fig. 4) elongate; surface generally smooth, shining but rough on depression of hypomeron for reception of pro-femur and tibia and on depressions of mesepimeron and metasternum for reception of midleg; punctures large and umbilicate, but on elytra small; depressions of hypomeron for reception of proleg and of mesepimeron and metasternum for reception of midleg without punctures; interspaces between punctures generally shorter than puncture diameter, but on metasternum, elytra and abdominal ventrites larger than puncture diameter and partly shorter than puncture diameter. Color. Body brown. Elytron with four sub-circular yellow spots, of which one spot medially on humerus, two spots side-by-side on anterior 1/3, and one spot medially on posterior 1/3. External edge of mandibles, mesal edge of antennal groove of hypomeron, sides of tarsal groove of pronotosternal sutures, external edge of scutellar shield, metasternal carina ventrad of depression for reception of mid-tarsi, striae and distal edge of elytron, and posterior edge of each abdominal ventrite blackish. Mouthparts yellowish with labrum and mandibles red-brown; antennae and legs orange with yellow tarsi. Body covered with yellow setae (Fig. 4). Head. Frontal margin broadly rounded in dorsal view (Fig. 5A); frons weakly depressed mesally (Fig. 5A, B); supra-antennal carina absent (Fig. 5B, C); frontoclypeal region most protruding between antennal sockets (Fig. 5D), grooved between eyes and mandibles (Fig. 5C); the grooves receive basal two antennomeres, opened posteriorly. Eyes 0.25–0.35 × longer than interocular distance in dorsal view. Mouthparts directed ventrally (Fig. 5D). Labrum elliptical, anterior edge more tapered (Fig. 5E), 0.5 × longer than wide. Antennae extended beyond half-length of pronotum and not reaching pronotal posterior lateral apices by antennomere XI (Fig. 4A, B); relative antennomere lengths: II<III<IV–X<XI<I (Fig. 5F); antennomere I elongate and curved cylindrical; II short cylindrical, 0.8– 0.85 × longer than wide; III trapezoidal, 1.1–1.4 × longer than wide, 2.0–2.4 × longer than II; IV–X parallelogram, weakly rounded distally, 1.1–1.3 × longer than wide; IV 1.1–1.2 × longer than III, 0.75–0.9 × longer than II–III combined; V 1.0–1.05 × longer than IV; XI bullet shaped, 1.7–2.1 × longer than wide, 1.3–1.5 × longer than X; each I–X with longitudinal carina distally; carinae on VI–X not reaching to their anterior edge. Mandibles unidentate (Fig. 5G); penicillus short, formed by brush of short setae; dorsal sinuous carina developed; with setae of various lengths on outside of the carina; outside concave basally (Fig. 5C). Maxilla (Fig. 5H): cardo constricted posteriorly; basistipe triangular; mediostipe rectangular; galea short, brush-like; lacinia short, brush-like; palpomeres short and broad; palpomeres II–III shorter than wide; palpomere IV fusiform, 1.2–1.3 × longer than wide. Labium (Fig. 5I): mentum wide trapezoidal, with long setae; prementum with cluster of short setae medially; apical palpomere subcylindrical. Prothorax widened posteriad on anterior half, slightly narrowed on posterior half ahead of posterior angles (Fig. 4A), widest on its half-length or across posterior lateral apices, broadly rounded laterally on posterior 2/3. Antennal grooves between pronotum and hypomeron deep and opened anteriorly, not reaching posterior lateral apices of prothorax, visible in lateral view (Fig. 5D); sides of antennal grooves carinate. Pronotum sub-hexagonal, 1.3 × longer than wide, roundly convex but weakly constricted posterior to anterior edge (Fig. 4A, B), tallest around the center (Fig. 5D), with median longitudinal depression posteriorly; each lateral carina extending from posterior angles along antennal grooves (Fig. 5D: black arrow), not reaching anterior angles; anterior angles acute (Fig. 5A); anterior edge straight in dorsal view (Fig. 5A); posterior angles extending posteriad, acute, with median longitudinal carina vestigial; posterior edge without sublateral incision near each hind angle, lobated and protruding medially. Hypomeron depressed for reception of pro-femur and tibia, carinate behind the depression; anterior angle sharply pointed (Fig. 5J: white arrow); posterior edge broadly rounded mesally and weakly emarginate near distal posterior angle; mesal posterior angle obtuse, not protruding (Fig. 5J: black arrow). Pronotosternal sutures (Fig. 4C) deeply grooved to receive pro-tarsi; tarsal groove opened posteriorly and connected to hypomeron depression for reception of pro-femur and tibia (Fig. 5J), becoming shallower anteriad, not reaching anterior edges of hypomeron and prosternum. Prosternum: anterior edge carinate, roundly projecting medially, depressed posterior to prosternal lobe; prosternal lobe short, not reaching the level of anterior edge of prothorax (Fig. 5D). Prosternal process 2.0–2.3 × longer than procoxal cavity length, depressed between procoxae, strongly curved dorsad behind procoxae (Fig. 5D); dorsal margin broadly rounded but with shallow concave medially in lateral view (Fig. 5D); ventral lobe almost parallel sided and gradually narrowed near apex in ventral view, with median longitudinal carina on posterior 1/3 (Fig. 4C); the carina forming pointed subapical tooth (Fig. 5D: white arrow); ventral margin rounded inward anteriorly and straight posteriorly in lateral view (Fig. 5D); ventral apex reaching dorsal apex; apex rounded in ventral and lateral views (Figs. 4C, 5D). Scutellar shield distinctly inclined anterior-downwards but posterior part parallel to elytral surface (Fig. 4A, B), tongue-shaped in anterodorsal view (Fig. 5K), narrowed posteriad but almost parallel-sided posteriorly, 1.1–1.3 × longer than wide, convex (Fig. 5L); anterior part strongly protruding anteriad (Figs. 4A, 5L); posterior part visible in lateral view (Fig. 5D: gray arrow); anterior angles rounded; anterior edge weakly and widely emarginated from anterior view; apex rounded or narrowly emarginate apically. Mesosternum (Fig. 6A): anterior edge straight but anterior edge of mesosternal cavity weakly protruding, lobate on each side; mesosternal cavity floor without median band of setae; mesosternal cavity almost parallel-sided anteriorly in ventral view; borders of mesosternal cavity in lateral view rounded anteriorly, straight medially and then curved ventrad obtusely (Fig. 5D); mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view (Fig. 5D); posterior edge 0.15–0.2 × wider than total width of mesosternum, almost straight (Fig. 6A); mesosternum and metasternum connate medially (Fig. 6A). Mesepisternum excavate anteriorly (Fig. 6A). Mesepimeron depressed for reception of mid-femur (Fig. 5J). Mesocoxal cavity closed to mesepisternum by mesosternum and mesepimeron (Fig. 5J). Elytron 5.2–5.45 × longer than wide, 2.0–2.2 × longer than pronotal length, parallel-sided but narrowed from posterior 1/3, rounded apically, broadly convex but plane medially (Fig. 4A), with striae. Elytral apices not meeting at midline (Fig. 4A, B). Hind wings (Fig. 6B) fully developed, without veins CuA 1 and CuA 3+4; RA 3+4 vestigial (Fig. 6B: white arrow); r4 translucent; mediomedial-cross vein between MP 1+2 and MP 3+4 vestigial (Fig. 6B: black arrow); mediocubital cross-vein between MP 3+4 and CuA 2 anteriad to contact point between MP 3 and MP 4, not complete (Fig. 6B: grey arrow); AA3 located at contact point between CuA 2 and CAS; radial cell 3.7–4.2 × longer than wide. Metepisternum narrower than width of elytral epipleuron (Fig. 5J). Metasternum depressed for reception of mid-tibia and tarsi, carinate ahead of depression for reception of mid-tibia and ventrad of depression for reception of mid-tarsi (Fig. 5J: grey arrow), sulcate medially and ahead of metacoxal cavities (Fig. 4C). Metacoxal plate narrowed medially; lateral edge distinctly longer than width of metepisternum; lateral anterior angle rounded; lateral posterior angle broadly pointed (Fig. 6C: arrow). Tibiae with a pair of simple spurs (Fig. 6D: arrow); relative tarsomere lengths: IV<III<II<V<I (Fig. 6D); tarsi and claws simple. Abdomen. Ventrite V curved triangular, rounded apically (Fig. 4C), 0.55–0.6 × longer than wide. Tergites and sternites VIII‒X yellow. Tergite VIII (Fig. 6E) longer than wide, almost paradell-sided basally but weakly sinuate, broadly rounded apically. Sternite VIII (Fig. 6F) triangular, abruptly narrowed at its half-length, shorter than wide; apex rounded. Tergite I