Dilobitarsus pendleburyi Fleutiaux 1934

Dilobitarsus pendleburyi Fleutiaux, 1934 (Figures 5–8) Dilobitarsus pendleburyi Fleutiaux, 1934: 178 (original description; type locality: Malaysia, Sabah, Bettotan, near Sandakan); Hayek, 1973: 99 (generic review; examination of holotype); Arimoto, 2016: 38 (redescription; recorded from the Malay Peninsula and Sumatra); Kasmiatun et al., 2020: 37 (recorded from Sumatra). Non-type material. 1 male, Malaysia, Pahang, Fraser’s Hill, 9 V 2010, Kiyoshi Matsuda leg. [DPF01; OMNH]; 1 female, Malaysia, Sarawak, Miri, Lambir Hills National Park, 23 VIII 2003, Keiko Kishimoto-Yamada & Takao Itioka leg. [AA1223; KIFDS]; 1 female, Indonesia, West Sumatra, near Payakumbuh, Harau Valley, 15 VIII 1992, A. Sarimudanas leg. [DPF02; OMNH]. Diagnosis. Dilobitarsus pendleburyi is distinguished from the congeners by the following combination of characters: pronotum with transverse black band formed by black scale-like setae ahead of posterior tubercles (Fig. 5A, B); elytra with white U-shaped band formed by white scale-like setae along apical margin (Fig. 5A, B); frontal carina V-shaped in ventral view (Fig. 6B, C); nasal plate high laterally (Fig. 6C); antennae extended beyond pronotal posterior apices by antennomere XI in male (Fig. 5A); antennomere III 1.3–1.6 x longer than wide (Fig. 6E, F); antennomeres IV–V 1.2–1.35 x longer than wide (Fig. 6E, F); antennomere XI 2.4–2.7 x longer than wide (Fig. 6E, F); pronotum with six tubercles (Fig. 6H: arrows); prosternal carina extending anterad of coxal cavities (Fig. 6I: arrow); anterior edge of mesosternum with small median notch (Fig. 6L); elytron broadly convex but abruptly depressed after strongly elevated tubercle (Fig. 7A); parameres with short lateral subapical barb (Fig. 7K, L); apex of parameres beyond lateral subapical barb 0.2 x total length of paramere (Fig. 7I, J), sub-trapezoidal and 1.5 x width at expansions in dorsal side (Fig. 7K); apical mesal edge of parameres emarginate between proximal and distal apices (Fig. 7K). Measurements. Male. BL: 11.6, BW: 3.05, MAE: 1.54, MBE: 0.93, OI: 165, PL: 3.62, PML: 2.98, PW: 2.89, PAW: 1.84, PLI: 125, PWI: 157, EL: 7.74, EW: 1.55, EI: 498, BI: 213. Female (2 specimens). BL: 8.99–15.3, BW: 2.39–3.98, MAE: 1.27–2.06, MBE: 0.78–1.25, OI: 162–164, PL: 2.89–4.75, PML: 2.52–4.05, PW: 2.28–3.78, PAW: 1.38–2.29, PLI: 126–127, PWI: 166, EL: 5.68–9.83, EW: 1.21–1.96, EI: 468–502, BI: 197–207. Additional description. Head. Eyes 0.3 x longer than interocular distance in dorsal view (Fig. 6A). Nasal plate protruding beyond base of labrum (Fig. 6D). Antennomere II 0.65–0.7 x longer than wide (Fig. 6E, F); III 1.3–1.6 x longer than wide, 2.5–3.2 x longer than II; IV 1.2–1.3 x longer than wide, 0.9–0.95 x longer than III, 0.65–0.7 x longer than II–III combined; V 1.2–1.35 x longer than wide, 0.9–1.1 x longer than IV; XI 2.4–2.7 x longer than wide, 1.5–1.7 x longer than X. Apical maxillary palpomere 1.8–1.85 x longer than wide (Fig. 6G). Thorax. Pronotum with six tubercles, of which anterior two elevated slightly (Fig. 6D: white arrow, 6H: white arrows) and posterior four elevated strongly (Fig. 6D: black arrows, 6H: black arrows), tallest at median posterior tubercles (Fig. 6D), without median longitudinal depression posteriorly; posterior edge without sublateral incision near each hind angle. Prosternal process 2.8–3.0 x longer than procoxal cavity length. Scutellar shield 1.2–1.3 x longer than wide, not visible in lateral view (Fig. 6D); anterior part inclined anterior-downwards and posterior part parallel to elytral surface (Fig. 6K). Mesosternum (Fig. 6L): anterior edge with small median notch, not lobate and sinuate on each side; mesosternal cavity almost parallel-sided anteriorly in ventral view; posterior edge 0.25–0.3 x wider than total width of mesosternum, straight (Fig. 6L). Elytron broadly convex but abruptly depressed after strongly elevated tubercle (Fig. 7A: arrow), with band on underside of tubercle (Fig. 7B: arrow). Hind wings (Fig. 7C) fully developed, without veins CuA 1 and CuA 3+4; r4 translucent; mediomedial cross-vein between MP 1+2 and MP 3+4 present; mediocubital cross-vein between MP 3+4 and CuA 2 vestigial; radial cell more than 4 x wide. Tarsomere IV with developed ventral lobes (Fig. 7D: arrow). Abdomen. Ventrite V curved triangular, rounded apically (Fig. 5C), 0.7–0.8 x longer than wide. Male. Tergites and sternites VIII‒X yellow (Fig. 7E‒H). Tergite IX (Fig. 7G); posterior median notch 0.2 x total length of tergite IX. Tergite X (Fig. 7G) almost as long as wide, 0.8 x length of tergite IX. Aedeagus yellow to brown (Fig. 7I, J). Phallobase 0.23 x total length of aedeagus, 0.72 x longer than wide. Median lobe; basal struts 0.2 x total length of median lobe. Parameres; apex beyond short lateral subapical barb 0.2 x total length of paramere, sub-trapezoidal and 1.5 x width at expansions in dorsal side (Fig. 7K); ventral side 1/2 x width of dorsal side around apex (Fig. 7L: arrow); proximal apex broadly rounded (Fig. 7K: white arrow); distal apex narrowly rounded (deformable by drying and other factors) (Fig. 7K: black arrow); apical mesal edge emarginate between proximal and distal apices. Female. Tergite VIII, sternite VIII and ovipositor yellow to brown (Fig. 8A–D); sclerotized plate inside bursa copulatrix orange (Fig. 8E). Tergite VIII (Fig. 8A) 1.3 x longer than wide. Sternite VIII length between base of spiculum ventrale and apex 0.7 x longer than wide (Fig. 8B). Spiculum ventrale 3.4 x longer than length of sternite VIII between base of spiculum ventrale and apex (Fig. 8C). Uterus with colleterial gland each on dorsal and ventral sides (Fig. 8F: arrows). Remarks. Arimoto (2016) described the pronotum as having four tubercles, but this was incorrect; actually, the pronotum has six tubercles. Distribution (Fig. 1: circles). Malaysia (Borneo, the Malay Peninsula), Indonesia (Sumatra). Ecology. This species was collected from both lowland and highland areas ~ 1,000 m in altitude, in May and August. The new specimen was collected using light traps.