Two New Species of Pachydactylus (Squamata: Gekkonidae) from the Kaokoveld

Recent surveys in southwestern Angola uncovered cryptic diversity within Pachydactylus caraculicus. Phylogenetic analyses of nuclear and mitochondrial DNA recover three distinct lineages. Two of these lineages, characterized by their typical banded dorsal pattern, show clear geographic structuring: a “northern” lineage corresponding to true P. caraculicus, and a distinct “southern” lineage extending into neighboring Namibia. The third lineage, sister to the clade of banded caraculicus, comprises morphologically divergent specimens resembling P. punctatus. All three lineages within the P. caraculicus complex are diagnosable based on morphological and coloration characters. We formally describe the two unnamed lineages of P. caraculicus as new species and discuss these findings in the biogeographic context of the Kaokoveld's lizard fauna.