Traumatomutilla incerta

Traumatomutilla incerta (Spinola, 1841) Figs 8–9 Mutilla incerta Spinola, 1841: 93. Mutilla dentata Smith, 1879: 219. Syn. nov. Mutilla sodalicia Kohl, 1882: 490. Syn. nov. Traumatomutilla weyrauchi Mickel, 1945: 30. Traumatomutilla dignitosa Mickel, 1952: 131. Syn. nov. Traumatomutilla tabatinga Casal, 1969: 287. Syn. nov. Ephuta (Traumatomutilla) sodalicia – André 1902: 56. Traumatomutilla sodalicia – André 1902: 40. Mutilla incerta – André 1902: 73 (incertae sedis). Traumatomutilla incerta – Mickel 1937: 196. Traumatomutilla dentata – Nonveiller 1990: 77. Diagnosis Female Occipital carina slightly but conspicuously swollen dorsolaterally; anterolateral carinae present in scutellar area, partially obscured by dense setation; lateral surface of propodeum predominantly smooth and shining, sculpture sparse; mesonotum with transverse band of silvery-white setae on posterior half; T2 with a pair of reddish to orange integumental spots. Male Apex of cuspis with short setae; pronotum densely clothed with appressed silvery-white setae; pronotal sculpture with dense micropunctures. Type material Lectotype of Mutilla incerta [FRENCH GUYANA] • ♀; Cayenne; D. Lebas leg.; MSNT. Holotype of Mutilla dentata BRAZIL • ♂; [Amazonas], St Paul [São Paulo de Olivença]; NHMUK. Holotype of Mutilla sodalicia BRAZIL • ♀; Amaz. [sic]; NHMW. Holotype of Traumatomutilla dignitosa GUIANA • ♂; Bartica, Bartica District; Wm. Beebe leg.; UMSP. Holotype of Traumatomutilla tabatinga BRAZIL • ♀; Amazonas, Tabatinga; Nov. 1958; F.M. Oliveira leg.; AMNH. Other material examined (68 ♀♀, 36 ♂♂) BOLIVIA – Beni • 1 ♀; Dpto. [Department], 5 km NW of Rurenabaque; 18 Sep. 1994; D. Roubik leg.; MIUP. – Cochabamba • 1 ♀; Region Chapare, 400 m [above sea level]; 20 Mar. 1907; Zischka; MNCN. – La Paz • 2 ♀♀; Yungas de Coroico; 1914; MNHN. – Locality unknown • 1 ♀; Tumupasu; 1921; W.M. Mann leg.; USNM. BRAZIL – Amapá • 1 ♀; Oiapoque, (Retiro) Ambiente de mata; 23 Oct. 2001; J. Madson leg.; HAMAB • 1 ♀; Serra Lombard, Limão; 26 Aug. 1961; J. and B. Bechyné leg.; MPEG. – Amazonas • 4 ♀♀; Manaus; RBINS • 1 ♀; Jul. 1962; F.M. Oliveira leg.; DZUP • 1 ♀; Manaus, Bairro Flores; 13–20 Apr. 2009; D. Storck-Tonon leg.; INPA • 1 ♀; Manaus, INPA 1 [Campus 1], Bosque da Ciência; 3°05′50″ S, 59°59′16″ W; 10 Jun. 2017; M.A. Bento leg.; INPA • 1 ♀; Manaus, Reserva Ducke, Trilha principal; 3 Oct. 2016; G. Amora leg.; INPA • 1 ♀; Manaus, Reserva Ducke; 2.93308° S, 58.97116° W; 19 May 2012; Medeiros leg.; INPA • 1 ♀; Manaus, Reserva Ducke; 2°55′48″ S, 59°58′31″ W; 8 Jun. 2013; J.F. Saraiva leg.; INPA • 1 ♀; Teffe [Tefé]; 5 Feb. 1920; H.S. Parish leg.; FMNH • 1 ♂; Tefé; 9 Sep. 1909; Ducke leg.; MNHN • 2 ♀♀; Tonantins, Amazon River; Jul. 1923; S.M. Klages leg.; CM • 2 ♀♀; Aug. 1923; S.M. Klages leg.; CM • 1 ♀; Manacapuru, Amazon River; Sep. 1923; S.M. Klages leg.; CM • 1 ♀; Estinao [Estirão do] Equador, Rio Javari; M. Alvarenga leg.; AMNH. – Mato Grosso • 1 ♂; Sinop; 12°31′ S, 55°37′ W; Oct. 1975; F.M. Oliveira leg.; USU. – Pará • 3 ♀♀; MZSP • 1 ♀; Óbidos; Dec. 1965; Maller, A. leg.; DZUP • 1 ♀; Mar. 1958; USNM • 1 ♀; USNM • 1 ♀; Jul. 1959; MNRJ • 2 ♀♀; Santarém; RBINS • 1 ♂; MNHN • 1 ♀; 1923; SDEI • 1 ♂; Altér do Chão; 12–13 Sep. 1991; J. Vidal leg.; INPA • 1 ♀; Oriximiná; 2 Sep. 1919; H.S. Parish leg.; FMNH • 2 ♂♂; Jacareacanga; Dec. 1968; M. Alvarenga leg.; AEIC • 8 ♂♂; Tucurui; Jan. 1979; M. Alvarenga leg.; AEIC. – Rondônia • 1 ♀; 62 km SE [kilometers southeast of] Ariquemes; 17–24 Mar. 1989; W.J. Hanson leg.; USU • 1 ♀; same data as for preceding; 5–16 Nov. 1996; W.J. Hanson leg.; USU • 1 ♂; Ouro Preto d’Oeste; 20 Sep. 1987; Elias, C. leg.; DZUP. COLOMBIA – Caquetá • 1 ♂; PNN [Parque Nacional Natural] Chiribiquete, Río Cuñare; 0°30′ N, 72°37′ W; 300 m [above sea leve]; 1–5 Jun. 2000; E. González and M. Ospina leg.; FSCA • 1 ♂; Río Cuñare-Amu; 0°13′ N, 72°25′ W; 300 m [above sea level]; 28 Feb.–3 Mar. 2001; M. Ospina and E. González leg.; USU. – Meta • 1 ♂; PNN [Parque Nacional Natural] Tinigua, Vda. [sic] Bajo Raudal; 2°16′ N, 73°48′ W; 460 m [above sea level]; 1–15 Mar. 2003; C. Sánchez leg.; IAvH. – Putumayo • 1 ♂; Mocoa; 4 Jul. 1978; M. Cooper leg.; NHMUK • 1 ♂; Santa Rosa, Kofan Indian village; 10 Oct. 1970; B. Malkin leg.; FMNH • 1 ♂; PNN [Parque Nacional Natural] La Paya, Resguardo Cecílio Cocha; 0°11′ S, 74°55′ W; 200 m [above sea level; 20–24 Jan. 2003; C. Sarmiento leg.; IAvH. – Locality unknown – 2 ♂♂; IAvH • 1 ♀; Rio Negro [sic], Ost Columb. [western Colombia]; MNCN ECUADOR – El Oro • 1 ♀; 21–25 km E [east of] Atahualpa; 27–31 Sep. 1997; F.T. Hovore leg.; UCDC • 1 ♀; 25 km E [east of] Atahualpa; 1°00′ S, 78°00′ W; 1–6 Oct. 1997; F.T. Hovore leg.; USU. – Morona Santiago • 1 ♀; Santiago; Feb. 1986; L. Coloma leg.; MIUP. – Napo • 1 ♀; Tiputini Biodiversity Station; 20 Jul. 1999; M.J. Miller leg.; UAIC • 1 ♀; Archidona; R. Haensch leg.; MNCN • 1 ♀; Misahualli, nr. [near] Tena; 10 Jan. 1989; B.J. Nichols leg.; UMRM • 1 ♀; Misahualli, nr. [near] Tena; 13–20 Jun. 1998; C. and K. Messenger leg.; UNSM • 1 ♀; same data as for preceding; 6–19 Oct. 2001; C. Brammer leg.; USU • 1 ♀; Río Aguarico; Mar. 1992; E.S. Ross leg.; CASC. – Sucumbios • 1 ♂; Sucumbios River, Sacha Lodge; 3–23 Jun. 1994; USU • 1 ♂; Río Napo, Sacha Lodge; 00°30′ S, 76°30′ W; 220–230 m [above sea level]; 27 Aug.–10 Sep. 1994; P. Hibbs leg.; USU • 1 ♂; same data as for preceding; 13 Jul. 1994; P. Hibbs leg.; USU • 1 ♂; same data as for preceding; 270 m [above sea leve]; 13–25 Feb. 1994; P. Hibbs leg.; USU. – Locality unknown • 2 ♀♀; MNCN. FRENCH GUIANA – Cayenne • 1 ♀; ZMUC • 1 ♀; Chavrein; 1914; R. Benoist leg.; MNHN • 1 ♀; Gourdonville; 1914; R. Benoist leg.; MNHN. – Saint-Laurent-du-Maroni • 1 ♀; Maroni, St [Saint] Laurent du Maroni; E. Le Moult; MNCN. GUYANA – Cuyuni-Mazaruni • 1 ♂; Bartica: 1 Jun. 1924; AMNH • 1 ♀; Kartabo; Jul–Aug. 1920; W.M. Wheeler; MCZ • 1 ♀; jct. [juncture] Cuyani/Mazaruni, Essequibo Rivers; 1927; R.E. Fuglestad leg.; EMEC. PERU – Huánuco • 1 ♀; Tingo Maria; 3 Jan. 1970; J.C. Schuster leg.; FSCA • 2 ♂♂; 23 Oct. 1946; Pallister et al. leg.; AMNH • 1 ♀; vic. [vicinity] Tingo Maria; 1–5 Jun. 1999; W. Hanson and S. Keller leg.; USU • 2 ♂♂; Monson [Monzón] Valley; 16 Nov. 1954; USU. – Junín • 1 ♀; Satipo; 18 Jul. 1939; W.F. Walsh Jr leg.; MCZ. – Lima • 1 ♀; Valle Chanchamayo; 1939; P. Vaquero leg.; UMSP • 2 ♀♀; 1914; MNHN • 2 ♀♀; S. Thamm leg.; ZMB • 1 ♂; 21 Jul. 1949; J.M. Schunke leg.; NHMUK. – Loreto • 1 ♀; Amazon Camp, Río Momón, nr [near] Iquitos; 1–10 Dec. 1982; E.S. Ross leg.; CASC. – Madre de Dios • 1 ♀; Pakitza; 23–28 Feb. 1992; B.V. Brown leg.; USNM • 2 ♀♀; Pakitza; Feb. 1992; R. Cambra leg.; MIUP • 1 ♀, 1 ♂; Pakitza; 30 Jun.–4 Jul. 1993; R. Cambra leg.; MIUP. – Piura • 1 ♀; Santo Domingo; NHMUK. SURINAME – Brokopondo • 1 ♂; Brownsberg Nature Park, Irene Val Trail; 4°95′ N, 55°18′ W; 400 m [above sea level]; 25 Aug.–2 Sep. 2007; G. Kung and A. Kreuter leg.; USU. VENEZUELA – Amazonas • 1 ♀; Alto Río Sialba; 1°43′10″ N, 64°31′50″ W; 24–25 Feb. 1989; 510 m [above sea level]; T. Lattke leg.; FSCA. Description Female BODY LENGTH. 13–20 mm. HEAD. Posterior margin almost straight. Occipital slightly but conspicuously swollen dorsolaterally. Vertex width 0.8× pronotal width. Eye almost circular, its length in frontal view almost equal to distance from its ventral margin to mandibular condyle. Head sculpture partially concealed by dense setation, densely and finely foveolate-punctate with sharp intervals where visible. Genal carina well defined. Mandible oblique, tapering slightly towards apex with small subapical tooth. Dorsal scrobal carina present, well defined, narrowly separated from antennal tubercles and lateral scrobal carina. Antennal tubercle coarsely and irregularly rugose. Flagellomere 1 2.6× pedicel length; flagellomere 2 1.9× pedicel length. MESOSOMA. Dorsal thoracic length 0.75 × maximum dorsal thoracic width. Mesosomal dorsum partially concealed by dense setation, densely and finely foveolate-punctate with sharp intervals where visible. Anterior surface of propodeum defined, short, almost as long as pronotal collar, finely striated longitudinally throughout with sparse interspersed fine punctures dorsad; dorsal surface rounded into anterior surface in lateral view. Humeral carina well defined, strongly projected dorsally, narrowly connected to conspicuously projected angulate epaulet; anterolateral corners of pronotum sharply angulate in dorsal view. Pronotal spiracle slightly projected from lateral margin of pronotum. Lateral surface of pronotum with sparse scattered punctures and dense micropunctures except at smooth conspicuous acute tubercle anteroventral in relation to pronotal spiracle; mesopleuron partially concealed by dense setation, densely micropunctate anteriorly, densely and coarsely foveolate-punctate on mesopleural ridge where visible; metapleuron completely concealed by dense setation, except small dorsal fourth unsculptured, smooth and shining. Lateral face of propodeum predominantly unsculptured, smooth and shining, with sparse scattered punctures. Ratios of width of humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles, 57: 84: 74:57: 58. Lateral margin of mesonotum conspicuously constricted anterior to propodeal spiracle, strongly diverging anterad, medially projected, into blunt process; post-mesonotal tubercle absent. Propodeal spiracle strongly projected from lateral margin of mesosoma; post-spiracular area indistinguishable. Scutellar scale present, as wide or wider than surrounding sculpture; anterolateral carinae present, vestigial, approximately twice as wide as scutellar scale; scutellar area with irregular intervals, not scabrous. Propodeum convex in lateral view, dorsal surface indistinguishable from posterior surface. METASOMA. Ratios of width of T1, width of T2 and length of T2, 34: 73: 66. Disc of T2 mostly concealed by dense setation, densely and co