Clistopyga laplatense Bordera & Palacio & Herrera-Flórez 2025, sp. nov.

18. Clistopyga laplatense Bordera & Palacio, sp. nov. (Figs 28, 29, 53, Appendix I) Diagnosis. Clistopyga laplatense may be distinguished from all other species of the C. calixtoi species group by the following combination of characters: genal orbits widely interrupted with brown or black (Figs 28A–D, 29A, 29F); pronotum extensively orange, with narrow dorsal lateral band and anterior ventral part, yellow cream (Fig. 28D, 29E); mesopleuron ventrally white cream (28A, 28D, 29A, 29E); metasoma with posterolateral well defined white cream marks on tergites I–VI, clearly separated from posterior rim (Figs 28A, 28F–H, 29A, 29G); epomia as a very short carina at the bottom of the submarginal depression of pronotum (Figs 28D, 29E); hind leg with femur 3.45–3.68× as long as high (Fig. 28J); submetapleural carina extended on anterior 0.55–0.8 of metapleuron (Fig. 28C, 29E). Additionally, female has metapleuron 2.15–2.2x as long as high (Fig. 28C); ovipositor relatively stout, evenly and slightly to moderately up-curved on distal 0.3–0.35 (Figs 28A, 28K), 1.95–2.1× as long as hind tibia; ovipositor sheath 1.65–1.175× as long as hind tibia. Male has mesosoma extensively marked with red or orange (Figs 29A, 29E–F) and gena in frontal view straight or slightly convex in profile, without any traces or longitudinal concavity posterior to mandible base (Fig. 29B). Description. Female: Body length 6.0– 8.8 mm. Fore wing length 4.1–6.0 mm. Head (Figs 28A–B, 28D). In dorsal view, moderately narrowed posteriorly to eyes. Gena shiny with very sparse setiferous punctures, in dorsal view slightly rounded, 0.33–0.35× as long as eye, in frontal view strongly constricted below eyes, slightly rounded and convex. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 0.9–1.25 its maximum diameter. Distance between hind ocelli 1.0–1.25× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform posteriorly to mandible. Face with shallow and moderately dense setiferous punctures, distance between them less than its diameter. Clypeal suture moderately curved. Clypeus 2.0–2.2× as broad as medially long, moderately convex with isolated fine punctures dorsally, flat and smooth ventrally, ventral margin slightly incurved. Malar space 0.9–1.0× as long as basal mandibular width, with shallow and slightly granulate subocular sulcus. Antenna with 24–27 flagellomeres, first flagellomere 5.3–5.75× as long as medially wide. Mesosoma (Figs 28A, 28C–E, 28G, 28I–J). Pronotum smooth and shiny with few setiferous punctures on posterior dorsal corner. Epomia as a very short carina at the bottom of the submarginal depression of pronotum. Mesoscutum shiny, with moderately dense very shallow setiferous punctures, denser on anterior part of median lobe; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus weak, reaching about 0.5 the length of mesoscutum. Mesopleuron shiny, with evenly sparse shallow setiferous punctures except at mid and dorsal posterior part. Epicnemial carina weak, its dorsal end straight, ending at level of centre of pronotum. Metapleuron, smooth and shiny, glabrous, 2.15–2.2× as long as high. Submetapleural carina extended on anterior 0.55–0.7 of metapleuron. Propodeum shiny with evenly sparse fine setiferous punctures, in dorsal view 1.05–1.15× as long as medially wide. Propodeal spiracle 0.13–0.16 the length to anterior end of pleural carina. Hind leg with femur 3.45–3.65× as long as high, 1.0–1.05× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m 0.3–0.4× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.3–1.5× as long as Cu 1 b. Hind wing with vein cu-a 0.3–0.35 × as long as abscissa of Cu 1 between M and cu-a, vein cu-a slightly reclivous, first abscissa of Cu 1 vertical, sometimes vein cu-a + first abscissa of Cu 1 not angled, vertical; distal abscissa of Cu 1 absent. Metasoma (Figs 28A, 28F, 28H, 28J). Tergite I 1.45–1.5× as long as posteriorly broad, shiny, with sparse shallow setiferous punctures laterally and posteriorly, spiracle near its anterior 0.35; lateromedian longitudinal carinae present at anterior 0.3 of tergite; lateral longitudinal carinae present at anterior 0.2 and posterior 0.3 of tergite. Sternite I extending posteriorly 0.50–0.55 the length of tergite. Tergite II 1.2–1.4× as long as posteriorly broad, with dense deep punctures, distance between punctures less than its diameter; anterior dorsal part of tergites III–VI granulate. Ovipositor relatively stout, evenly up-curved at 0.3–0.35 distal, 1.95–2.1× as long as hind tibia. Ovipositor sheath 1.65–1.75× as long as hind tibia, ventral margin basally without denticles, length of setae on average 2.5–3.0× the sheath basal width. Colouration (Fig. 28). Head mostly white cream, with apex of mandibles, lateral parts of clypeus, fovea, wide longitudinal stripe at middle of face, frons and vertex medially, middle and dorsal part of gena and occipital region, dark brown to black; antenna brown, scapus, pedicel, annellus and first to second flagellomere lateroventrally, white cream. Mesosoma mostly orange; dorsal part of propleuron, brown; propodeum dorsally dark orange, black posteriorly; propleuron ventrally, dorsal lateral band and anterior part of pronotum, two longitudinal stripes on mesoscutum posteriorly to notauli, tegula, subalar prominence, scutellum dorsally, postscutellum, wide diffuse longitudinal mark on ventral part of mesopleuron, mesepimeron, and lateral parts of propodeum, yellow cream. Metasoma brown; tergites I–VI with white cream posterolateral marks, tergites II–V with dark brown to black posterior band; tergite I with yellowish anterior lateral marks; ovipositor brown; ovipositor sheaths black. Legs predominantly white cream; front leg with dorsal stripes on femur and tibia, and distal part of tarsi, brown; mid leg with base of trochanter, dorsal and ventral stripes on femur and tibia, and distal part of tarsi, brown; hind leg with anterior and posterior spots on coxa, base of trochanter, base, and anterior and posterior subdistal marks on femur, basal and distal part of tibia, and distal part of tarsi, brown. Wings hyaline, pterostigma blackish brown. Male: Body length 5.2–8.0 mm. Fore wing length 3.4–4.5 mm. Head (Figs 29A–B, 29E–F). Gena in dorsal view 0.43–0.45× as long as eye, in frontal view strongly constricted ventrally to eyes, slightly rounded and convex. Posterior ocellus separated from eye 0.9–1.0 its maximum diameter. Distance between hind ocelli 0.9–1.0× maximum diameter of posterior ocellus. Clypeus about 1.8× as broad as medially long. Antenna with 23–26 flagellomeres, first flagellomere 5.0–5.25× as long as medially wide. Mesosoma (Figs 29A, 29C–F). Epicnemial carina weak, its dorsal end slightly towards the back. Metapleuron, smooth and shiny, glabrous, 1.9–1.95× as long as high. Submetapleural carina strong extended on anterior 0.7–0.8 of metapleuron. Propodeum in dorsal view 1.0–1.1× as long as medially wide. Hind leg with femur 3.67–3.68× as long as high, 0.86× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m about 0.3× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a about 1.3× as long as Cu 1 b. Hind wing with vein cu-a + abscissa of Cu 1 between M and cu-a not angled, vertical. Metasoma (Figs 29A, 29G). Tergite I 1.75–1.9× as long as posteriorly broad, shiny, with shallow moderately dense punctures laterally and posteriorly, spiracle near its anterior 0.4; lateral longitudinal carinae present at posterior 0.2 of tergite. Sternite I extending posteriorly 0.55 the length of tergite. Tergite II 1.25–1.3× as long as posteriorly broad, with dense and deep punctures, distance between punctures less than its diameter; anterior dorsal part of tergites III–VI not granulate. Colouration (Fig. 29). As in female but without central brown mark on face. Other characters as in female. Etymology. The specific name refers to the city of La Plata (Argentina) and surrounding localities, where most of the type material was collected. Type material (17 ♀♀, 4 ♂♂). Holotype. Argentina: 1 ♀, Berisso, 8.XIII.1965 H. & M. Townes (EMUS). Paratypes. Argentina: 1 ♀, Berisso, 8-XII-1965, H. & M. Townes (EMUS); 1 ♀, La Plata, 15-XII-1965, H. & M. Townes (EMUS); 1 ♀, same data, 16-XII-1965 (EMUS); 1 ♀, 1 ♂, same data, 24-I-1966 (EMUS); 1 ♀, same data, 25-I-1966 (EMUS); 1 ♀, same data, 27-I-1966 (EMUS); 3 ♀♀, same data, 29-I-1966 (EMUS); 1 ♀, 2 ♂♂, same data, 30-I-1966 (EMUS); 1 ♀, same data, 31-I-1966 (EMUS); 1 ♀, Punta Lara, 26-I-1966, H. & M. Townes (EMUS); 1 ♂, Tacanas, Tucumán, 5–30-X-1968, Lionel Stange (EMUS). Brazil: 1 ♀, Nova Teutonia, 27º11'B 52º23'L, 26-IV-1938, Fritz Plaumann, B.M. 1938-682 (NHMUK); 1 ♀, same data, 17.XI.1938, B.M. 1938-312 (NHMUK); 1 ♀, Nova Teutonia, Santa Catarina, 20-XII-1951, Fritz Plaumann (EMUS); 1 ♀, Rondon, 24º38'B 54º07'L, 6-X-1952, Fritz Plaumann (EMUS). Distribution. Argentina, Brazil (Fig. 53, Appendix I).