Cicindela coerulea Pallas 1773

Cicindela (s. str.) coerulea nitida Lichtenstein, 1796 Figs 10–13, 19–23, 45–54, 76–82, 89–132 Cicindela nitida Lichtenstein, 1796: 32 (Type locality—Sibiria). = Cicindela tricolor M.F. Adams, 1817: 278 (Type locality—Sibiria ad lacum Baikal et ad ripas fl. Lenae, ultra urbem Jakutzk), synonymized by Horn (1915: 337). = Cicindela tricolor var. obliquefasciata Fischer von Waldheim, 1828: 21, Tab. 1 fig. 3 (Type locality—Sibiriae), synonymized by Roeschke (in Horn & Roeschke 1891: 49). = Cicindela tricolor var. tenuifascia Fischer von Waldheim, 1828: 22, Tab. 1 fig. 6 (Type locality— Altai), synonymized by Roeschke (in Horn & Roeschke 1891: 49). = Cicindela optata Fischer von Waldheim, 1828: 22, Tab. 1 fig. 8 (Type locality—Dauria, ad litora fluvii Amur), synonymized by Motschoulski (1844: 30). = Cicindela tricolor var. saltatoria Gistl, 1837: 38 (Type locality—Sibiria ad lacum Baikal et ad ripas fl. Lenae, ultra urbem Jakutzk, et in montosis Altaï), synonymized by Horn (1915: 337). = Cicindela coerulea selengensis Mandl, 1936: 12 (Type locality—Selengatal, Selengafe), synonymized by Schilder (1953: 564). Cicindela tricolor M.F. Adams, 1817: Fischer von Waldheim 1820: 6, Tab. 1 fig. 3, Tab. 17 fig. 3; 1821: 102; 1828: 12, 21; 1829: 5; Dejean 1833: 3; 1836: 3; Dejean & Boisduval 1837: 24 Pl. 3 fig. 3; Sturm 1843: 2; Motschoulski, 1844: 30; Motschulsky, 1850: 1; Chaudoir 1865: 36; Sahlberg 1880: 5; Fleutiaux 1892: 106; Horn & Roeschke 1891: 48, Taf. V fig. 11; Tschitschérine, 1903: 94; Jakobson 1905: Tab. 1 fig. 3; Horn 1905a: 157, 158; 1905b: 27; Poppius 1906: 14; Lafer 1978: 5, figs 5, 14; 1989: 96, fig. 52.4, 97. Cicindela hybrida tricolor M.F. Adams, 1817: Jakobson 1905: 190; Horn 1905a: 157, 158. Cicindela optata Fischer von Waldheim, 1828: Motschulsky, 1850: 1. Cicindela nitida Lichtenstein, 1796: Lutshnik 1916: 95; Mandl 1921: 79; Shilenkov 1994: 8; Kryzhanovskij et al. 1995: 26; Berlov & Berlov 1997: 44; Sundukov 1998: 9; Matalin 2000: 676, fig. 35; Rogatnykh 2006: 30; Rogatnykh & Yakubovich 2009: 108. Cicindela hybrida nitida Lichtenstein, 1796: Mandl 1921: 80, 81; Winkler 1924: 4; Horn 1915: 337; 1926: 221; 1930: 402; Wu 1937: 15; Institute of Zoology Academia Sinica 1986: 417, fig. 11-1-3; Hu 1992: 374; Zhu et al. 1999: 26, 27 Tab. 4 fig. 13; Fang & Wu 2001: 69. Cicindela grumi nitida Lichtenstein, 1796: Schilder 1953: 564. Cicindela nitida tricolor M.F. Adams, 1817: Shilenkov 1987: 7. Cicindela nitida optata Fischer von Waldheim, 1828: Berlov & Berlov 1997: 44. Cicindela nitida selengensis Mandl, 1936: Kryzhanovskij et al. 1995: 27; Matalin 2000: 676, fig. 34. Cicindela coerulea Pallas, 1773: Rogatnykh 2009a: 9; 2009b: 69; Sundukov 2009: 89; Khobrakova 2012: 167; Putchkov 2013: 54 fig. 2.4, 62, 63, 65 fig. 4.1, 68 fig. 7.4, 73 fig. 12.5–6, 80. Cicindela coerulea nitida Lichtenstein, 1796: Lutshnik 1924: 21; Okninskyi 1930: 20; Mandl, 1935: 285 fig 1b, 294 figs 1–5, 296 figs 1–5; 1936: 11; 1965: 241; 1974: 137; Horn 1938: 46, Taf. 63 figs 23–25; Nakane 1960: 30, 38 Pl. I fig. d; Mandl 1970: 73 Abb. 3 fig. 6, 74; Alekseeva 1977: 5; Cassola & van Nidek 1984: 10; Werner, 1991: 18, T. 14 figs 102, 104–106, T. 15 figs 107–109; Li 1992: 5; Wiesner 1992: 117; 2020: 183; Putchkov & Shilenkov 1996: 451; Lorenz 1998: 42; 2005: 131; Puchkov & Matalin 2003: 104; Khobrakova & Sharova 2004: 144; Shook & Wiesner 2006: 10; Khobrakova & Dudko 2010: 37; Wu & Shook 2010: 78; Wu 2011: 24; Budilov 2012: 40; Sundukov 2013: 76; Khobrakova et al. 2014: 70; Koshkin et al. 2016: 312; Matalin & Dudko 2016a: 140; 2016b: 143; Tsuji et al. 2016: 3; Putchkov & Matalin 2017: 227; Matalin et al. 2019: 10; National Species List of Korea 2019: 199; Makarov et al. 2020; Putchkov 2020: 169 fig. 45.1, 173 fig. 49.4, 182 fig. 56.5, 199 fig. 61.1; Shilenkov 2021: 41. Cicindela coerulea selengensis Mandl, 1936: Mandl, 1935: 294 figs 6–8, 296 figs 6–10, 1936: 12; Horn 1938: 46, Taf. 63 fig. 26; Cassola & van Nidek 1984: 10; Werner, 1991: 18, T. 15 figs 112, 113; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 43; 2005: 131; Puchkov & Matalin 2003: 104; Putchkov & Matalin 2017: 227. Type material. Lectotype of Cicindela optata Fischer von Waldheim, 1828 (designated here), ♂ (Figs 45, 76, 77)— C. optata Fisch., Nertshinsk [handwritten white label], Typus [typed pink label], Fischer von Waldheim [typed white label], Staatl. Museum für Tierkunde Dresden [typed white label], Lectotypus, Cicindela (s. str.) nitida f. optata F.-W., design. A. V. Matalin, 1997 [handwritten and typed red label]; LECTOTYPUS, Cicindela optata Fischer von Waldheim, 1828, design. A. V. Matalin, 2024 [printed red label] (MTD). Paralectotype of Cicindela optata Fischer von Waldheim, 1828 (designated here), ♂ —Staatl. Museum für Tierkunde Dresden [typed white label], Paralectotypus, Cicindela (s. str.) nitida f. optata F.-W., design. A. V. Matalin, 1997 [handwritten and typed red label], PARALECTOTYPUS, Cicindela optata Fischer von Waldheim, 1828, design. A. V. Matalin, 2024 [printed red label] (MTD). Paratypes of Cicindela coerulea selengensis Mandl: 1♂ (Fig. 113)— Transbaic., Selenga fl. [handwritten white label], Сollect. Hauser [typed white label], C. tricolor var. [handwritten white label], v. optata, det. Ganglbauer [typed and handwritten white label], Cicindela nitida Licht. s. selengensis m., det. Ing. Mandl [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label]; Cicindela (s. str.) coerulea (Pallas) selengensis Mandl 1936, Det. J. Probst [printed white label], NHMW-ZOO-COL-0002631 [printed white label]; 1♀ —Transbaic., Selenga fl. [handwritten white label], Сollect. Hauser [typed white label], v. optata, det. Ganglbauer [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002632 [printed white label]; 1♀ (Fig. 114)— ibid, NHMW-ZOO-COL-0002633 [printed white label]; 1♀ (Fig. 112)—Transbaicalia, Selenga Tal., V – VI.1908 [typed white label], Сollect. Hauser [typed white label], v. optata, det. Ganglbauer [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002634 [printed white label]; 1♀ —Transbaic., Selenga fl. [handwritten white label], Сollect. Hauser” [typed white label], v. optata, det. Ganglbauer [typed and handwritten white label], Cicindela nitida Licht. s. selengensis m., det. Ing. Mandl [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002635 [printed white label]; 1♂ (Figs 80, 81)— ibid, NHMW-ZOO-COL-0002638 [printed white label]; 1♀ —Cotype [typed red label], Selenga-Tal, Transbaikalien [handwritten white label], Cicindela nitida Licht. ssp. selengensis m., det. Ing. Mandl [typed and handwritten white label], NHMW-Acquisitions-Nr.: Col-1971-6, coll. E. Grundmann [printed white label], NHMW-ZOO-COL-0002639 [printed white label]. Additional material. RUSSIA, Altai Republic: 1♂ —Chuysky trakt, Ust-Inya, 15.VII.1907, E.G. Rodd.; 1♀ — Naryn-Gol River, 1956, Pridanieva; 1♀ — Altai, env. Ongudai, leg. A.G. Yakobson (all ZIN); 3♀♀ — ibid, 2– 3. VI.1989, S. Saluk; 1♀ —Chemal, 30.VII.[19]63; 1♂ —mouth of Chulcha River, right tributary of Chulyshman River, 29.VIII.[19]34, V. Geptner; 1♂ —mouth of Malyi Karakem River, tributary of Chulyshman River, 25. VI.1989, leg. V.N. Dremin (all ZMUM); 1♂ — Altai, Seminskiy pass, VII.1979; 2♂♂ 2♀♀ —valley of Chuya River, ZhanaAul, 1850 m, 8.VII.2022, leg. A. et E. Filippov (both MSPU); 3♂♂ 1♀ —Altaigeb., Akash, Mittellauf des Mjonka, Kuraikette, 1600–2300 m, 12.6.1992 (cJW); 2 ex. — Altai, Akash, Mjonka-Fluss, 20.05.1993, leg. A. Budarin; 9 ex. — ibid, 14.06.1991, leg. O. Legezin (both cJG); 1♀ —SE Altai, Kurai, 1.VIII.[1]964, Kerzhner; 1♂ 1♀ —Buguzun River, Chikhacheva Mnt Ridge, ~1900 m, 29.VII.[1]964, Grunin; 1♂ 1♀ —Chagan-Uzun env., bank of Chuya River, 5–6. VI.1989, V. Grichik et S. Saluk; 1♀ — ibid, A.S. Pisanenko (all ZIN); Republic of Khakassia: 2♂♂ 2♀♀ —Shira Lake, 5.VIII.1916, Khvorov; (ZMUM); 2♂♂ — Tomsk Gouv., Fluss Abakan, Ui-Bat River, 17.VII., A. Jakobson; 1♂ 1♀ — Tomsk Gouv., Tashtyb, 16. VI., A. Jakobson (both ZIN); 1 ex. —Abakan, Krasnoe Lake, 26.07.1993, leg. A. V. Chuvilin (cFB); 2♂♂ 1♀ —Beisk District, near Tabat vill., 19.VIII.[19]63, O. Grigorjev (ISEA); 3♂♂ 3♀♀ —2–3 km S of Abakan, 5.VII.1997, leg. V. Patrikeev; 1♂ —valley of Son River, 2. VI.1969, leg. V. Patrusheva (both MSPU); 3♀♀ —Borets, bank of Son River, 54.46 oN 90.36 oE, h~380 m, 29.06.2011, leg. K. Tomkovich (cOM); Republic of Tuva: 6♂♂ 9♀♀ —Erzin, sands on the shore of Tere-Khol, 7. VI.1962, leg. D. Berman (ZMUM); 1 ex. — Tuva, Erzin, Erzin-River, 30.05.1990, leg. S. Gundorov (cJG); 2♀♀ —17 km NE of Khadyn Lake, 26. V.1988, leg. V. Zinchenko; 1♀ —Balgazyn, 8. VI.1948, Cherepanov (both ISEA); 1♂ — ibid, 28.VIII.1959, leg. S. V. Sharova (cISl); 1♀ —Balgazyn, 110 km SE of Kyzyl, VII.1956, Galkin; 6♂♂ —Toore-Khem, 11.08.1973, leg. B. Korotyaev; 1♂ —near Erzin, 21.07.1988, leg. S. Byzov (all ZIN); 1♂ 3♀♀ —SE shore of ToreKhol Lake, 31. V –1. VI; 1♂ —Kyzyl District, bank of Biy-Khem River, near Kara-Khaak, 20. V.1997, leg. V. Patrikeev; 2♂♂ 3♀♀ —Shagonar, 26.07.1992, A. Napolov; 2♂♂ —near Kyzyl, bank of Enisey River, 1–6.VII.1997, leg. S. Vastchenko & R. Mishustin; 6♂♂ —Khadyn Lake, 16–19. VI.1997, leg. S. Vastchenko & R. Mishustin; (all MSPU); 1♂ — ibid, 25. V.1998, leg. A. Gorodinski (cXX); 1 ex. — ibid, 20.05.1998, leg. S. Vastchenko (cFB); 1♂ 1♀ — ibid, 25. VI.1998; 2♂♂ —30 km N Balgazyn, 21.6.1994, leg. P. Smrz, Z. Kletecka (both cJW); Krasnoyarsky Krai: 1♂ —Bazaikha near of Krasnoyarsk, 14. V.[19]01, Krutovsky; 1♂ — ibid, 9. VI.[19]01; 19♂♂ 20♀♀ —Kaptyrevo; 10♂♂ — Krasnoyarsk (all ZIN); 1♂ 1♀ —O Sibiria, Berezovka, leg. Dr. J. Fleischer (cJW); 3♂ 5♀ —Sibiria, Minusinsk, Exp. Zichy, leg. Csiki (HNHM); 1♂ —Minusinsk, Opytnoe Pole, 18. V.1928; 5♂♂ 4♀♀ — ibid, 25– 31.VII.1928 (both ISEA); 1♂ —Sibiria, Krasnoyarsk, A. Zolo