Cephennodes (Fusionodes) spatulatus Jałoszyński, 2025, sp. nov.

Cephennodes (Fusionodes) spatulatus sp. nov. (Figs 206–209) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “CHINA: Yunnan, mountain W / Gejiu, 23º24’13’’N, 103º07’28’’E, / 1990 m, mixed forest, litter and / various debris sifted, 25.VIII.2014, / leg. M. Schülke [CH14-20 b]” [white, printed]; “ CEPHENNODES / (FUSIONODES) / spatulatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (2 exx.): 2 ♀♀, same data as for holotype, each with yellow PARATYPUS labels (cPJ, MNB). Diagnosis. Body moderately large, BL around 1.5–1.6 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser, punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 207–209) in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta. Description. Body of male (Fig. 206) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.55 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 mm, HW 0.33 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.73 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 about as long as broad, 4–10 each transverse, 11 about as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, equally broad between base and slightly anterior to middle; PL 0.48 mm, PW 0.65 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i.e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra distinctly wider than pronotum, together oval, broadest near anterior third; EL 0.88 mm, EW 0.73 mm, EI 1.21; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings not studied. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 207–209) AeL 0.40 mm; in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle, their outer lateral margins strongly sinuate; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta. Female. Externally indistinguishable from male, wings not studied; BL 1.51–1.60 mm; HL 0.21–0.23 mm, HW 0.33–0.35 mm, AnL 0.68–0.75 mm; PL 0.48–0.50 mm, PW 0.70 mm; EL 0.83–0.88 mm, EW 0.73 mm, EI 1.14–1.21. Distribution. S China: Yunnan. Etymology. The adjective spatulatus refers to the spatulate shape of the dorsal subapical projection of the aedeagus in lateral view. Remarks. Cephennodes spatulatus is the only member of the C. hippopotamus species group that has the aedeagus in lateral view with two conspicuously massive dorsal subapical projections, of which distal one is strongly broadening towards apex, and proximal one is much shorter and distally tapering.

Publication Details

Journal
Zenodo (CERN European Organization for Nuclear Research)
Published
2025-07-18
DOI
https://doi.org/10.5281/zenodo.16606720
Primary Topic
Subterranean biodiversity and taxonomy
Type
article
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article

Cephennodes (Fusionodes) spatulatus Jałoszyński, 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Subterranean biodiversity and taxonomy
article

Cephennodes (Fusionodes) spatulatus Jałoszyński, 2025, sp. nov.

article en

Abstract

Cephennodes (Fusionodes) spatulatus sp. nov. (Figs 206–209) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “CHINA: Yunnan, mountain W / Gejiu, 23º24’13’’N, 103º07’28’’E, / 1990 m, mixed forest, litter and / various debris sifted, 25.VIII.2014, / leg. M. Schülke [CH14-20 b]” [white, printed]; “ CEPHENNODES / (FUSIONODES) / spatulatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (2 exx.): 2 ♀♀, same data as for holotype, each with yellow PARATYPUS labels (cPJ, MNB). Diagnosis. Body moderately large, BL around 1.5–1.6 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser, punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 207–209) in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta. Description. Body of male (Fig. 206) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.55 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 mm, HW 0.33 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.73 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 about as long as broad, 4–10 each transverse, 11 about as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, equally broad between base and slightly anterior to middle; PL 0.48 mm, PW 0.65 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i.e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra distinctly wider than pronotum, together oval, broadest near anterior third; EL 0.88 mm, EW 0.73 mm, EI 1.21; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings not studied. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 207–209) AeL 0.40 mm; in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle, their outer lateral margins strongly sinuate; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta. Female. Externally indistinguishable from male, wings not studied; BL 1.51–1.60 mm; HL 0.21–0.23 mm, HW 0.33–0.35 mm, AnL 0.68–0.75 mm; PL 0.48–0.50 mm, PW 0.70 mm; EL 0.83–0.88 mm, EW 0.73 mm, EI 1.14–1.21. Distribution. S China: Yunnan. Etymology. The adjective spatulatus refers to the spatulate shape of the dorsal subapical projection of the aedeagus in lateral view. Remarks. Cephennodes spatulatus is the only member of the C. hippopotamus species group that has the aedeagus in lateral view with two conspicuously massive dorsal subapical projections, of which distal one is strongly broadening towards apex, and proximal one is much shorter and distally tapering.

Zenodo (CERN European Organization for Nuclear Research)
Life in Land
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Subterranean biodiversity and taxonomy
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