Tritia reticulata

Tritia reticulata (Linnaeus, 1758) (Fig. 12A 1-A 3) Buccinum reticulatum Linnaeus, 1758: 740. Nassa (Hinia) reticulata – Harmer 1925: 318, pl. 34, figs 1, 2. Hinia (Hinia) reticulata – Cuerda Barceló 1987: 295, pl. 27, fig. 19. Nassarius reticulatus – Landau et al. 2009: 29, pl. 5, figs 15-18, pl. 6, fig. 1, pl. 17, fig. 5. — Chirli & Linse 2011: 160, pl. 54, fig. 3. — Rijken & Pouwer 2014: 54, fig. 35. — Van Dingenen et al. 2015: pl. 8, fig. 1. For more, see synonymy list in Landau et al. (2009) and Van Dingenen et al. (2015). MATERIAL AND DIMENSIONS. — Maximum height 24.3 mm, width 12.3 mm. — RGM.1364925 (2), leg. WG; RGM.1365204 (1), leg. ACJ; RGM.1365007 (4), leg. ACJ; RGM.1310337 (1), leg. AWJ; RGM.1365111 (40), leg. AWJ. SPECIES CHARACTERISATION. — Shell solid, bucciniform, with elevated conical spire. Protoconch not preserved. Teleoconch of five weakly convex whorls, separated by a shallowly impressed suture. Sculpture of low axial ribs overrun by strap-like cords swollen over the intersection giving surface reticulated appearance. Aperture pyriform; outer lip thickened by varix, denticulated within; columella bearing two folds abapically and parietal fold adapically; columellar callus thickened; parietal callus extended over venter forming parietal shield. DISTRIBUTION. — Lower Pliocene: Atlantic, SE Spain (González-Delgado 1989; Landau et al. 2011); western Mediterranean, NE Spain (Gili 1991), SE France (Fontannes 1879, Cossmann 1901); central Mediterranean, Italy (Bellardi 1882; Chirli 2000), Tunisia (Fekih 1975). — Upper Pliocene: Atlantic, Portugal (Zbyszewski 1959; Gili et al. 1995; Silva 2001); western Mediterranean, S Spain (Landau et al. 2009); central Mediterranean, Italy (Bellardi 1882; Malatesta 1974). — Upper Pliocene-Lower Pleistocene: Atlantic, NW France (Van Dingenen et al. 2015). — Lower Pleistocene: Atlantic, Selsoif, NW France (Van Dingenen et al. 2015); central Mediterranean, Italy (Cerulli-Irelli 1911); eastern Mediterranean, Rhodes Island (Chirli & Linse 2011). — Middle Pleistocene: Atlantic, Wexford Gravels, Irish Sea (Harmer 1916). — Pleistocene (indeterminate): Netherlands (Rijken & Pouwer 2014). — Upper Pleistocene: western Mediterranean, Balearic Islands (Cuerda Barceló 1987). — Holocene: NSB (Strand Petersen 2004). Today, it occurs in the Atlantic and western Mediterranean (Rolán & Luque 1994). REMARKS Within the Tritia reticulata species group (of Landau et al. 2009), T. nitida (Jeffreys, 1867) differs in having a paucispiral protoconch of less than two whorls, T. reticulata has 2.75-3.25 whorls. Moreover, T. nitida has a slightly gradate spire, fewer axial ribs and stronger and more regular dentition within the outer lip. Specimens from Selsoif show the same variability in axial sculpture as seen in other populations. For further discussion see Landau et al. (2009: 30).