Inella pseudolongissima Rolan & Fernandez-Garces 2008

“ Inella ” pseudolongissima Rolán & Fernández-Garcés, 2008 Fig. 6 Inella pseudolongissima Rolán & Fernández-Garcés, 2008: 102, fig. 11. “ Inella ” pseudolongissima – Fernandes & Pimenta 2019a: fig. 3n. Type material Holotype CUBA • sh; off Havana; 823 m (as stated in the original description) or 444–823 m (as stated on the NMNH invertebrate collection website); Blake Expedition; USNM 87316. Paratypes See Rolán & Fernández-Garcés (2008). Material examined GUADELOUPE (Karubenthos 2 expedition) • 5 sh; stn DW4549; MNHN • 4 sh; stn DW4550; MNHN • 2 sh; stn DW4589; MNHN • 1 sh juv.; stn DW4599; MNHN • 1 sh; stn DW4601; MNHN • 1 sh; stn DW4634; MNHN. Emended description Shell sinistral, conical-fusiform, up to 19.9 mm long (adult shells reach at least 12.6 mm in length), 2.7 mm wide, length/width ratio 6.2–8.0, apical angle of early whorls 11–13°. Protoconch multispiral, columnar, 4.0–6.0 whorls, 0.84–1.48 mm long, 0.55–0.66 mm wide; first whorl smooth, not inflated; on second whorl two thin spiral cordlets appear (situated at 34–37% and 66–73% of whorl height), but abapical spiral cordlet sometimes thinner than adapical one and may appear only on third whorl; both cordlets may resemble only slight elevations in some shells instead of true cordlets due to their minute width; transition to teleoconch gradual, nearly indistinct. Teleoconch with up to 28 whorls; two spiral cords (adapical and abapical) in beginning of teleoconch, continuous with cordlets of protoconch; median spiral cord emerges narrowly between fourth and sixth whorls, bordering closely to adapical cord, slowly developing and reaching nearly same size as other cords only on body whorl (abapical cord can be slightly more developed than other cords in late whorls); suture shallow, with a smooth sutural cord; 18–19 opisthocline axial ribs on 18 th teleoconch whorl; medium-sized, nearly rounded to slightly elliptical nodules; slightly nodulous to nearly smooth subperipheral cord, with a smooth, thin basal cord right below it; no supranumerical cord; elliptical aperture, 1.4–1.5 mm long, 1.0– 1.3 mm wide, length/ width ratio 1.2–1.3; anterior canal open to partially open, directed downward, 0.60–0.74 mm long, 0.46–0.53 mm wide, length/width ratio 1.3–1.5; posterior canal as a deep sinus, almost detached from aperture. Shell white to faintly cream. Remarks The current record from Guadeloupe extends the known range of “ I.” pseudolongissima by ~ 2300 km. Agreeing with the original description and illustrations, shells from Guadeloupe are highly variable with respect to the size and number of protoconch whorls (although always multispiral, with at least four whorls), the strength of the spiral cordlets of the protoconch and the strength of the abapical spiral cord of the teleoconch (Fig. 6). The variation in the size and number of protoconch whorls may be related to the duration of embryonal growth within the egg; see Fernandes & Pimenta (2019a) for a discussion on other triphorids with a supposed long duration of intracapsular metamorphosis, resulting in multispiral protoconchs for species with a supposed non-planktotrophic development. Geographic distribution USA: Florida (Rolán & Fernández-Garcés 2008); Cuba (Rolán & Fernández-Garcés 2008); Guadeloupe (this study). Bathymetric distribution Empty shells previously known from 77–823 m (Rolán & Fernández-Garcés 2008). This study: 150– 632 m (empty shells).