Diochus

Key to species of the genus Diochus of Bhutan, India, Nepal, Myanmar, Pakistan and Thailand (Figs 1−13) 1 Elytra uniformly colored, dark brown to yellowish brown..................................................... 2 - Elytra with basal part black, and apical part mostly reddish yellow (Fig. 13).......... Diochus bicoloripennis Coiffait, 1981 2 Antennomeres long, third antennomere 3−4 times longer than wide.............................................. 3 - Antennomeres short, third antennomere 1−2.5 times longer than wide............................................ 5 3 Elytra brown, distinctly lighter than pronotum; posterior margin of male sternite VIII emarginate medially, with numerous short setae in emargination (Figs 5-1E−L); aedeagus with a single hooked asymmetrical sclerite on top (Figs 5-1I−M)................................................................................................ D. setatus sp. nov. - Elytra dark brown to black brown, about the same color as pronotum; posterior margin of male sternite VIII emarginate and either without setae in emargination or with numerous longer setae along entire apex; aedeagus with internal sclerites all symmetrical......................................................................................... 4 4 Pronotum elliptical, anterior region near anterior angles distinctly deflexed and narrowed; posterior margin of male sternite VIII emarginate medially, with numerous setae (Figs 9-1B−C, H); aedeagus with paired horn-like sclerites, parameres shorter (Figs 9-2A−D); spermatheca in basal part with widened tube (Figs 9-2A−D)................ D. conicollis (Motschulsky, 1858) - Pronotum not elliptical; anterior region near anterior angles not so deflexed and narrowed; posterior margin of male sternite VIII emarginate medially, without setae in emargination (Figs 2-1E, J); aedeagus without paired horn-like sclerites, parameres longer (Figs 2-2 A−D); spermatheca in basal part with tube not widened (Figs 2-2 E−H).............. D. cameroni sp. nov. 5 Fore body reddish ochraceous; tempora markedly widened posteriad; lateral margins of pronotum nearly parallel (Figs 11 A−B; antennae shorter, antennomeres 6−10 slightly transverse................................ D. ochraceus Cameron, 1926 - Fore body dark brown to yellowish brown; lateral margins of pronotum slightly widened posteriad; antennae longer, antennomeres 6−10 not transverse........................................................................ 6 6 Anterior region of pronotum near anterior angles deflexed, lateral margins distinctly widened posteriad................. 7 - Anterior region of pronotum near anterior angles slightly deflexed, lateral margins slightly widened posteriad or nearly parallel............................................................................................. 8 7 Head elliptical (Figs 7-1A, E); male sternite VIII widely and shallowly emarginate medially with almost straight median part (Figs 7-1C, H); aedeagus elongate, slightly widened apically, parameres very short (Figs 7-1D, L−M). Female unknown...................................................................................... D. ceylonicus sp. nov. - Head rounded oblong (Fig 8-1A); male sternite VIII triangularly emarginate medially; aedeagus elongate, narrowed apically, parameres longer (Figs 8-1I−J)....................................... D. guizhouensis Shuai, Nozaki & Tang, 2021 8 Head markedly elongate (Figs 6-1 A, D); male sternite VIII widely, shallowly and triangularly emarginate; aedeagus with paired horn-like sclerites and helical structures (Figs 6-2A−D); spermatheca with basal part of tube narrow (Figs 6-2E−F)...................................................................................... D. keralanus sp. nov. - Head slightly elongate; male sternite VIII, aedeagus and spermatheca different..................................... 9 9 Body small, head without elevation between eyes in male (Figs 12-1A, E); male sternite VIII very long, slightly emarginate apically (Figs 12-1C−D, I), aedeagus with transverse sclerite on top and irregular sclerites in middle (Figs 12-2A−D), spermatheca apically with a loop (Figs 12-2E−G)................................. D. antennatus (Motschulsky, 1858) - Body large, head with elevation between eyes in male; male sternite VIII shorter; aedeagus without transverse sclerite, spermatheca different, without circle apically.............................................................. 10 10 Posterior margin of male sternite VIII without setae, roundly emarginate (Figs 10-1C−D), aedeagus narrowed apically, parameres longer nearly as long as median lobe (Figs 10-1E−G); spermatheca with short tube, with only one bend (Figs 10-2C−E)........................................................................... D. punctipennis (Motschulsky, 1858) - Posterior margin of male sternite VIII with setae; aedeagus not or less narrowed apically, parameres shorter, much shorter than median lobe; spermatheca different, tube longer, with more bends.............................................. 11 11 Posterior margin of male sternite VIII with fascicular setae on both sides (Fig 3-1E); aedeagus with parameres thicker and shorter (Figs 3-1 I−J)................................................................. D. emarginatus sp. nov. - Posterior margin of male sternite VIII with only normal setae on both sides (Fig 4-1E); aedeagus with parameres thinner and longer (Figs 4-1I−J)...................................................................... D. setifer sp. nov.

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Publication Details

Journal
Zenodo (CERN European Organization for Nuclear Research)
Published
2024-11-20
DOI
https://doi.org/10.5281/zenodo.14248193
Primary Topic
Fish Biology and Ecology Studies
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article
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Diochus

Zenodo (CERN European Organization for Nuclear Research)
Fish Biology and Ecology Studies
article

Diochus

article en

Abstract

Key to species of the genus Diochus of Bhutan, India, Nepal, Myanmar, Pakistan and Thailand (Figs 1−13) 1 Elytra uniformly colored, dark brown to yellowish brown..................................................... 2 - Elytra with basal part black, and apical part mostly reddish yellow (Fig. 13).......... Diochus bicoloripennis Coiffait, 1981 2 Antennomeres long, third antennomere 3−4 times longer than wide.............................................. 3 - Antennomeres short, third antennomere 1−2.5 times longer than wide............................................ 5 3 Elytra brown, distinctly lighter than pronotum; posterior margin of male sternite VIII emarginate medially, with numerous short setae in emargination (Figs 5-1E−L); aedeagus with a single hooked asymmetrical sclerite on top (Figs 5-1I−M)................................................................................................ D. setatus sp. nov. - Elytra dark brown to black brown, about the same color as pronotum; posterior margin of male sternite VIII emarginate and either without setae in emargination or with numerous longer setae along entire apex; aedeagus with internal sclerites all symmetrical......................................................................................... 4 4 Pronotum elliptical, anterior region near anterior angles distinctly deflexed and narrowed; posterior margin of male sternite VIII emarginate medially, with numerous setae (Figs 9-1B−C, H); aedeagus with paired horn-like sclerites, parameres shorter (Figs 9-2A−D); spermatheca in basal part with widened tube (Figs 9-2A−D)................ D. conicollis (Motschulsky, 1858) - Pronotum not elliptical; anterior region near anterior angles not so deflexed and narrowed; posterior margin of male sternite VIII emarginate medially, without setae in emargination (Figs 2-1E, J); aedeagus without paired horn-like sclerites, parameres longer (Figs 2-2 A−D); spermatheca in basal part with tube not widened (Figs 2-2 E−H).............. D. cameroni sp. nov. 5 Fore body reddish ochraceous; tempora markedly widened posteriad; lateral margins of pronotum nearly parallel (Figs 11 A−B; antennae shorter, antennomeres 6−10 slightly transverse................................ D. ochraceus Cameron, 1926 - Fore body dark brown to yellowish brown; lateral margins of pronotum slightly widened posteriad; antennae longer, antennomeres 6−10 not transverse........................................................................ 6 6 Anterior region of pronotum near anterior angles deflexed, lateral margins distinctly widened posteriad................. 7 - Anterior region of pronotum near anterior angles slightly deflexed, lateral margins slightly widened posteriad or nearly parallel............................................................................................. 8 7 Head elliptical (Figs 7-1A, E); male sternite VIII widely and shallowly emarginate medially with almost straight median part (Figs 7-1C, H); aedeagus elongate, slightly widened apically, parameres very short (Figs 7-1D, L−M). Female unknown...................................................................................... D. ceylonicus sp. nov. - Head rounded oblong (Fig 8-1A); male sternite VIII triangularly emarginate medially; aedeagus elongate, narrowed apically, parameres longer (Figs 8-1I−J)....................................... D. guizhouensis Shuai, Nozaki & Tang, 2021 8 Head markedly elongate (Figs 6-1 A, D); male sternite VIII widely, shallowly and triangularly emarginate; aedeagus with paired horn-like sclerites and helical structures (Figs 6-2A−D); spermatheca with basal part of tube narrow (Figs 6-2E−F)...................................................................................... D. keralanus sp. nov. - Head slightly elongate; male sternite VIII, aedeagus and spermatheca different..................................... 9 9 Body small, head without elevation between eyes in male (Figs 12-1A, E); male sternite VIII very long, slightly emarginate apically (Figs 12-1C−D, I), aedeagus with transverse sclerite on top and irregular sclerites in middle (Figs 12-2A−D), spermatheca apically with a loop (Figs 12-2E−G)................................. D. antennatus (Motschulsky, 1858) - Body large, head with elevation between eyes in male; male sternite VIII shorter; aedeagus without transverse sclerite, spermatheca different, without circle apically.............................................................. 10 10 Posterior margin of male sternite VIII without setae, roundly emarginate (Figs 10-1C−D), aedeagus narrowed apically, parameres longer nearly as long as median lobe (Figs 10-1E−G); spermatheca with short tube, with only one bend (Figs 10-2C−E)........................................................................... D. punctipennis (Motschulsky, 1858) - Posterior margin of male sternite VIII with setae; aedeagus not or less narrowed apically, parameres shorter, much shorter than median lobe; spermatheca different, tube longer, with more bends.............................................. 11 11 Posterior margin of male sternite VIII with fascicular setae on both sides (Fig 3-1E); aedeagus with parameres thicker and shorter (Figs 3-1 I−J)................................................................. D. emarginatus sp. nov. - Posterior margin of male sternite VIII with only normal setae on both sides (Fig 4-1E); aedeagus with parameres thinner and longer (Figs 4-1I−J)...................................................................... D. setifer sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Institute of Zoology (CN)
Openalex Percentile: Top 40%
Fish Biology and Ecology Studies
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