Portunoidea Rafinesque 1815

Superfamily Portunoidea Rafinesque, 1815 Remarks: While the taxonomy of Portunoidea has previously been well studied, the phylogenetic relationships between the various groups now in the superfamily have been less settled with different systems being proposed (e.g., see like Schubart and Reuschel (2009) and Spiridonov et al. (2014). N. Evans (2018), based on morphological and molecular evidence, only recognized Geryonidae Colosi, 1924, Carcinidae MacLeay, 1838 and Portunidae Rafinesque, 1815 under the Portunoidea, leaving the status of one family, Brusiniidae Števčić, 1991 unresolved (but see Spiridonov 2020). We retain the Brusiniidae in the Portunoidea until more studies can be done. We follow Spiridonov et al. (2014) in recognizing the Podophthalminae Dana, 1851 and Lupocyclinae Alcock, 1899 as subfamilies under the Portunidae, and elevating Monomia Gistel, 1848 and Xiphonectes A. Milne-Edwards, 1873, to full genera; and N. Evans (2018) in recognizing Goniosupradens Davie & Ng, 2024 as full genus, and Thranita Evans, 2018 for members of the “ Thalamita prymna group”. In the Portuninae, the genera Portunus, Monomia and Xiphonectes had been shown to be composite (N. Evans 2018), many were resolved by Koch et al. (2022). For taxa treated herein, several species previously placed under Xiphonectes have recently been placed in new genera, a scheme we also follow.