Onthophagus landolti Harold 1880

Onthophagus landolti Harold, 1880 Fig. 60A Onthophagus landolti Harold, 1880: 34 (original description). Type locality: Colombia: Norte de Santander: Ocaña; and Venezuela: La Guaira: La Guaira. Name-bearing type: syntypes (MNHN), examined by FZVM. Onthophagus landolti – Bates 1877: 75 (distribution). — Schaeffer 1914: 293, 299 (key, note). — Boucomont & Gillet 1927: 206 (catalogue).— Boucomont 1932: 314 (synopsis, key). — Blackwelder 1944: 211 (checklist). — Roze 1955: 46 (checklist for Venezuela). — Howden & Cartwright 1963: 8, 94 (list, comments). — Howden & Yound 1981: 112 (contribution). — Gillet 1911b: 206 (catalogue). — Deloya 1992: 22 (list). — Halffter et al. 1992: 141, 148 (ecology). — Escobar 1997: 422–423 (diversity, list); 2000: 209 (checklist for Colombia). — Zunino & Halffter 1997: 162 (list). — Maes 1998: 655 (list for Nicaragua). — Kohlmann & Solís 2001: 214 (revision for Costa Rica). — Medina et al. 2001: 139 (checklist for Colombia). — Navarrete-Heredia et al. 2001: 57 (list). — Estrada & Coates-Estrada 2002: 1911 (ecology). — Bustos-Gómez & Lopera-Toro 2003: 61 (diet). — Hernández et al. 2003: 96 (diversity). — Zunino 2003: 71‒72 (cited for Mexico). — Krajcik 2006: 114 (checklist). — Kohlmann et al. 2007: 32 (atlas). — Pulido-Herrera & Zunino 2007: 107 (synopsis). — Pulido-Herrera et al. 2007: 307 (cited for Andean region of Colombia). — Lozano 2010: 86 (list). — Cultid-Medina et al. 2012: 54 (guide). — Delgado et al. 2012: 327 (list). — Ferrer-Paris et al. 2013: 109. — Delgado & Curoe 2014: 64 (key, cited for Panama). — Pérez-Cogollo et al. 2017: 119 (biology). — Nieto et al. 2020: 136 (report). — Moctezuma 2021: 186 (list, cited for Mexico). Onthophagus landolti landolti – Pulido-Herrera & Zunino 2007: 107 (cited as junior synonym). — Moctezuma 2021: 186 (list, cited for Mexico). Distribution United States, Mexico, Belize, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia and Venezuela (Pulido-Herrera & Zunino 2007). Subregions of Venezuela Maracaibo Depression, Plains, Central Coast Mountain Range, and Cerro Copey, Margarita Island. Literature records Harold 1880: 34 (Venezuela: La Guaira). — Bates 1886–1890: 75 (Venezuela). — Boucomont 1932: 326 (Venezuela). — Blackwelder 1944: 211 (Venezuela). — Roze 1955: 46 (Venezuela: Nueva Esparta: Isla Margarita; and Guárico). — Kohlmann & Solís 2001: 214 (Venezuela). — Zunino 2003: 71 (Venezuela). — Pulido-Herrera & Zunino 2007: 107 (Venezuela: La Guaira). — Lozano 2010: 86 (Venezuela: Zulia). — Ferrer-Paris et al. 2013: 109 (Venezuela: Aragua [Guárico]:Altagracia de Orituco; and Zulia: Rosario de Perijá). — Pérez-Cogollo et al. 2017: 119 (Venezuela).