Coprophanaeus lancifer

Coprophanaeus lancifer (Linnaeus, 1767) Fig. 24A Scarabaeus lancifer Linnaeus, 1767: 544 (original description). Type locality: Brazil: Pará: Belém, Parque Estadual do Utinga, 01°25′07.2″ S, 48°25′47.6″ W. Name-bearing type: neotype (BMNH ‒ ex CEMT), designated by Maldaner et al. (2017), examined by MC and FZVM. Phanaeus heros Castelnau, 1840: 80 (original description). Type locality: French Guiana. Name-bearing type: unknown syntypes, likely destroyed (Maldaner et al. 2017). Phanaeus miles Castelnau, 1840: 80 (original description). Type locality: French Guiana. Name-bearing type: unknown syntypes, likely destroyed (Maldaner et al. 2017). Phanaeus septentrionalis Pessôa, 1934: 291 (original description). Type locality: Brazil: Pará: Belém. Name-bearing type: lectotype (MZSP), designated by Maldaner et al. (2017), not examined. Scarabaeus lancifer – Lichtenstein 1796: 5 (cited). — Nevinson 1892: 5 (list). — Arnaud 2002b: 23 (citation). Phanaeus lancifer – Erichson 1848: 563 (catalogue). — Bodkin 1919: 213 (notes). — Gillet 1911b: 84 (catalogue). — d’Olsoufieff 1924: 76 (distribution). Phanaeus heros – Nevinson 1892: 5 (cited). — Maldaner et al. 2017: 89 (cited). Phanaeus miles – Nevinson 1892: 5 (cited). — Maldaner et al. 2017: 89 (cited). Megaphanaeus lancifer – Blackwelder 1944: 209 (checklist). Phanaeus septentrionalis – Martínez & Pereira 1967: 60 (synonym of Phanaeus lancifer). — Arnaud 2002b: 23 (cited as synonym of Coprophanaeus lancifer). — Maldaner et al. 2017: 89 (types). Coprophanaeus (Megaphanaeus) lancifer – Edmonds 1972: 841 (comparative morphology). — Escobar 2000: 207 (checklist for Colombia). — Feer 2000: 32 (list); 2008: 62 (ecology); 2013: 767 (list for French Guiana). — Vaz-de-Mello 2000: 192 (checklist for Brazil). — Vítolo 2000: 599 (key for Colombia). — Medina et al. 2001: 140 (checklist for Colombia). — Feer & Pincebourde 2005: 30 (list). — Quintero & Roslin 2005: appendix A (ecology). — Hamel-Leigue et al. 2009: 54 (distribution for Bolivia). — Edmonds & Zídek 2010: 14, 16 (key, revision). — Brûlé et al. 2011a: 193 (inventory). — Larsen 2011: 98 (list for Suriname). — Vaz-de-Mello et al. 2011b: 86 (list); 2013: 96 (list for Suriname). — Ratcliffe 2013: 494 (ecology). — Bicknell et al. 2014: supp. file. — Silva et al. 2014: 348 (diversity). — Feer & Boissier 2015: 170 (list). — Pacheco & Vaz-de-Mello 2015: 2 (cited for Roraima State, Brazil). — Boilly et al. 2016: 87, 90 (key, catalogue for Guyane). — Maldaner et al. 2017: 89 (cited). — Hielkema & Hielkema 2019: 96 (catalogue for the Guianas). Distribution Venezuela, Guyana, Suriname, French Guiana, Brazil, Peru, and Bolivia. Subregions of Venezuela Peneplain of the Casiquiare River –Upper Orinoco, and System of hills and low piedmont mountains of the Guiana Shield. Literature record Edmonds & Zídek 2010: 14, 16 (Venezuela: Amazonas and Bolívar).