Ontherus sulcator

Ontherus sulcator (Fabricius, 1775) Fig. 58B Scarabaeus sulcator Fabricius, 1775: 27 (original description). Type locality: French Guiana. Name-bearing type: according to Zimsen (1964), three syntypes in ZMUK, but deemed lost without further explanation by Génier (1996). We ourselves have made no attempt to locate them, and so they may well be in ZMUK as Zimsen said. Copris nisus Castelnau, 1840: 79 (original description). Type locality: French Guiana. Name-bearing type: lectotype (MVMA), designated by Génier (1996), not examined. Copris ovalipennis Blanchard, 1846: 180 (original description). Type locality: Argentina: Corrientes. Name-bearing type: lectotype (MNHN), designated by Génier (1996), not examined. Copris senegalensis Gillet, 1910: 15 (original description). Type locality: Unknown. Originally said to be Senegal (“ Sénégal ”), but this is incorrect (Gillet 1911a). Name-bearing type: syntypes (RBINS) (Gillet 1910), not examined. Scarabaeus sulcator – Olivier 1789: 142 (redescription). — Fabricius 1792: 61 (redescription). Copris sulcator – Fabricius 1801: 53 (redescription). — Olivier, 1790: 168 (redescription). — Martínez 1959: 71 (cited). Copris nisus – Blanchard, 1846: 180 (comments). — Burmeister 1874: 126 (list for Argentina). — Martínez 1959: 71 (cited as synonym of Ontherus sulcator). Ontherus sulcator – Erichson 1847b: 761 (comments). — Harold 1869c: 59 (comments); 1869d: 1008 (catalogue). — Burmeister 1874: 126 (list for Argentina). — Gillet 1911a: 319 (list); 1911b: 58 (catalogue). — Bruch 1915: 541 (list). — Lucas 1920: 459 (catalogue). — Luederwaldt 1930: 106 (comments); 1931a: 371, 385, 389, 392, 408, 411, 416 (key, synonymy). — Pessôa & Lane 1941: 456, 458 (key). — Blackwelder 1944: 206 (checklist). — Martínez 1959: 71 (catalogue). — Vulcano & Pereira, 1967: 583 (key). — Forsyth & Gill 1993: 70 (as O. suclator [sic], cited for Guyana). — Escobar 2000: 209 (list). — Vaz-de-Mello 2000: 194 (checklist for Brazil). — Medina et al. 2001: 139 (checklist for Colombia). — Hamel-Leigue et al. 2006: 16 (inventory, cited for Bolivia). — Feer 2008: 62 (ecology, list). — Larsen et al. 2008: 1294 (list). — Krajcik 2012: 174 (checklist, cited for Cayenne). — Boilly & Vaz-de-Mello 2013: 112 (Fig. 43, key). — Ferrer-Paris et al. 2013: 109 (list). — Ratcliffe et al. 2015: 197 (checklist, cited for Peru). Copris ovalipennis – Harold 1869a: 503 (comments); 1869c: 59 (comments). — Burmeister 1874: 126 (list for Argentina). — Luederwaldt 1931a: 386 (cited as synonym of Ontherus nisus). — Martínez 1959: 71 (cited as synonym of Ontherus sulcator). Ontherus nisus – Harold 1869d: 1008 (catalogue, cited as synonym of O. sulcator). — Gillet 1911b: 58 (catalog, cited as synonym of O. sulcator). — Luederwaldt 1931a: 387, 392, 399, 400, 409, 411, 417 (redescription, distribution, key). — Pessôa & Lane 1941: 457, 458 (key, redescription). — Blackwelder 1944: 206 (cited as synonym of O. sulcator). — Lange 1947: 310 (list). — Martínez 1959: 71 (cited as synonym of O. sulcator). Pinotus ovalipennis – Harold 1869d: 1008 (catalogue, cited for Argentina: Corrientes). Ontherus ovalipennis – Gillet 1911b: 58 (catalog, cited as synonym of O. sulcator). — Luederwaldt 1931a: 387 (cited as synonym of O. nisus). — Pessôa & Lane 1941: 458 (cited as synonym of O. nisus). — Blackwelder 1944: 206 (cited as synonym of O. sulcator). — Martínez 1959: 71 (cited as synonym of O. sulcator). Copris senegalensis – Gillet 1911a: 319 (cited as synonym of O. sulcator); 1911b: 58 (catalog, cited as synonym of O. sulcator). — Bruch 1915: 541 (list, cited as synonym of O. sulcator). — Martínez 1959: 72 (cited as synonym of O. sulcator). Ontherus sulcator – Luederwaldt 1931a: 364 (type species). Ontherus senegalensis – Luederwaldt 1931a: 386, 387 (cited as synonym of Ontherus nisus). — Pessôa & Lane 1941: 458 (cited as synonym of O. nisus). — Blackwelder 1944: 206 (cited as synonym of O. sulcator). Ontherus (Ontherus) sulcator – Génier 1996: 74‒76 (revision). — Hielkema & Hielkema 2019: 57 (catalogue for the Guianas). Distribution Colombia, Venezuela, Trinidad and Tobago, Guyana, Suriname, French Guiana, Brazil, Ecuador, Peru, Bolivia, Paraguay, Argentina, and Uruguay (Génier 1996). Subregions of Venezuela System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní of Northeast Guyana Shield, and Guiana Shield. Literature records Luederwaldt 1931a: 387, 409, 417 (Venezuela). — Pessôa & Lane 1941: 457–458 (Venezuela). — Vulcano & Pereira 1967: 583 (Venezuela). — Génier 1996: 74 (Venezuela: Bolívar). — Larsen et al. 2008: 1294 (Venezuela: Bolívar). — Ferrer-Paris et al. 2013: 109 (Venezuela: Bolívar: Isla de Anacoco).