Triepeolus segregatus PCYU

Triepeolus segregatus (Cockerell, 1900) Figs 12N, 15E, 20B, 29 Epeolus occidentalis var. segregatus Cockerell, 1900: 361 (♀, ♂). Epeolus permixtus Cockerell, 1923: 94 (♀, ♂), syn. nov. Triepeolus brunnescens Cockerell & Sandhouse, 1924: 313 (♂), syn. nov. Triepeolus pacis Cockerell, 1925b: 201 (♂), syn. nov. Epeolus sarothrinus Cockerell, 1929: 103 (♀, ♂), syn. nov. Epeolus sarothrinus var. confluens Cockerell, 1929: 103 (♂), syn. nov. Triepeolus segregatus – Cockerell 1904: 38; 1907: 63 (redescription). Doeringiella (Triepeolus) sarothrinus – Vergara & Ayala 2002: 24 (in regional checklist). Proposed common name Segregated triepeolus. Diagnosis The following morphological features in combination tell T. segregatus apart from all other Triepeolus in the simplex species group: the axillae do not have pale pubescence all along their margins and are small to intermediate in size, each with the tip extending little if at all beyond the midlength of the mesoscutellum and the free portion distinctly pointed but mesally unattached to the mesoscutellum for less than 2/5 the medial length of the axilla (Figs 20B, 29B); the mesopleura laterally do not have erect/suberect, simple setae or have only sparse, short (<¼ MOD), erect/suberect, simple setae in addition to the usual appressed, branched setae (Fig. 29A, C); the T1 discal patch is trapezoidal with an inverted U- or horseshoe-shaped anteromedial projection, with the basal band so deeply invaginated that it appears to be completely interrupted medially in dorsal view (Figs 15E, 29B); and the T2 apical transverse band has a pair of well-defined basomedially convergent anterolateral extensions (Figs 15E, 29A–C). Specimens from Texas and adjacent Mexico may be confused with T. parahirsutus sp. nov. and specimens from California with T. saturninus due to their similar size and trapezoidal discal patch with an inverted U- or horseshoe-shaped anteromedial projection, but in the two similar-looking species the mesopleura have sparse, pale-yellow, erect/suberect, simple setae (reaching about ½ MOD in length) in addition to the usual appressed, branched setae. Etymology The specific epithet is a Latin adjective meaning ‘set apart’, presumably in reference to Cockerell’s (1900) recognition of the holotype as a separate variety that “[d]iffers from typical [E.] occidentalis from Colorado ”, with which it was mistakenly considered to be conspecific. Material examined Primary type material MEXICO – Baja California • ♂, E. permixtus holotype; Isla Pond bay, Isla Ángel de la Guarda, Gulf of California; 1 Jul. 1921; E.P. Van Duzee leg.; CAS 954. – Baja California Sur • ♂, T. pacis holotype; La Paz; 29 Jun. 1919; G.F. Ferris leg.; CAS 1661. USA – California • ♂, T. brunnescens holotype; Poway, San Diego County; 10 Sep. 1884; F.E. Blaisdell leg.; CAS 1606 • ♀, E. sarothrinus holotype; Riverside, Riverside County; 26 May 1928; P.H. Timberlake leg.; AMNH AMNH_IZC 00323550 • ♂, E. sarothrinus var. confluens holotype; same data as for preceding; 26 May 1928; P.H. Timberlake leg.; USNM 534623. – New Mexico • ♂, E. occidentalis var. segregatus holotype; Las Vegas Hot Springs; 11 Jul.; USNM 534868. Secondary type material USA • 1 ♀, E. sarothrinus paratype; California, Riverside, Riverside County; 6 Jul. 1928; P.H. Timberlake leg.; AMNH M.G.R. Database No. 1516. DNA barcoded material Available. Three BINs in two clusters: Cluster: BOLD:AAG7909+BOLD:AFP1028. BIN: BOLD:AAG7909. Specimen examined and sequenced: USA • 1 ♀; California, Chollas Creek Park (San Diego), San Diego County; 32.7201° N, 117.0780° W; 15 Jul. 2011; K.L.J. Hung leg.; BOLD sample ID: KJH.4621; UCR. Cluster: BOLD:AAG7909+BOLD:AFP1028. BIN: BOLD:AFP1028. Specimens examined and sequenced: USA – Arizona • 1 ♀ (previously assigned to the BIN BOLD: AAG7909); Sierra Vista, Cochise County; 31.5100° N, 110.2590° W; 4 Sep. 2009; C.S. Sheffield and S. Dumesh leg.; BOLD sample ID: CMNTO_025; PCYU PCYU-SD09:631 • 1 ♂, T. cf. segregatus (previously assigned to the BIN BOLD: AAG7909); Southwestern Research Station (5 mi W of Portal), Cochise County; 22 Aug. 2007; H.T. Ngo and J. Gibbs leg.; BOLD sample ID: CMNTO_485; PCYU • 2 ♂♂, T. cf. segregatus (BIN previously unassigned); same data as for preceding; 22 Aug. 2007; H.T. Ngo and J. Gibbs leg.; BOLD sample IDs: CMNTO_487, CMNTO_488; PCYU. Cluster: BOLD:AAG7909+BOLD:AFP1028. BIN unassigned: USA – Arizona • 1 ♂, T. cf. segregatus; Southwestern Research Station (5 mi W of Portal), Cochise County; 22 Aug. 2007; H.T. Ngo and J. Gibbs leg.; BOLD sample ID: CMNTO_489; PCYU • 1 ♂; Southwestern Research Station (5 mi W of Portal), Cochise County; 31.8830° N, 109.2070° W; 21 Aug. 2019; J. Gibbs leg.; BOLD sample ID: WRME0501356; WRME WRME0501356. Cluster and BIN: BOLD:AEI6013. Specimens examined and sequenced: USA – Arizona • 1 ♂; 19 mi SE of Willcox, Cochise County; 18–25 Aug. 2002; BOLD sample ID: CCDB-38770 F09; RSKM RSKM_ENT_E-168248 • 1 ♂; Catalina Hwy (Santa Catalina Mountains), Coronado National Forest; 32.3631° N, 110.7137° W; 16 Aug. 2015; T.M. Onuferko leg.; BOLD sample ID: CMNTO_047; TOPC • 1 ♂; Catalina Hwy (Santa Catalina Mountains), Coronado National Forest; 32.3728° N, 110.7068° W; 16–26 Aug. 2015; T.M. Onuferko leg.; BOLD sample ID: CMNTO_109; PCYU • 1 ♀; Southwestern Research Station (5 mi W of Portal), Cochise County; 1 Sep. 2013; BOLD sample ID: CMNTO_017; PCYU • 1 ♀; Southwestern Research Station (5 mi W of Portal), Cochise County; 31.8840° N, 109.2060° W; 25 Aug. 2011; N. de Silva leg.; BOLD sample ID: 14511G09- AZ; PCYU 0003776 • 1 ♀; Southwestern Research Station (5 mi W of Portal), Cochise County; 31.8852° N, 109.2051° W; 24 Aug. 2017; R. Oram leg.; BOLD sample ID: CCDB-38770 F11; RSKM RSKM_ENT_E-167432. – Colorado • 1 ♀; Escalante State Wildlife Area, Sawmill Mesa Road, Delta County; 38.7361° N, 108.1556° W; 31 Aug. 2013; M. Buck leg.; BOLD sample ID: CMNTO_314; RAM pmae00122773. – New Mexico • 1 ♀; 7 km NE of Silver City, Grant County; 32.8270° N, 108.2410° W; 2 Sep. 2009; C.S. Sheffield and S. Dumesh leg.; BOLD sample ID: CMNTO_019; PCYU PCYU-SD09:289 • 1 ♀; 28 mi S of Animas, Hidalgo County; 31.5550° N, 108.8760° W; 27 Aug. 2019; J. Gibbs leg.; BOLD sample ID: CMNTO_162; WRME WRME0500765 • 1 ♀; NM-152 (along Iron Creek), Gila National Forest, Grant County; 32.9078° N, 107.8160° W; 18 Aug. 2015; T.M. Onuferko leg.; BOLD sample ID: CMNTO_050; TOPC. Non-barcoded material MEXICO – Baja California • 1 ♀; 1 mi E of Santa Inés; 6 Apr. 1985; Bloomfield and Faulkner leg.; INHS 387489 • 1 ♂; 3.3 mi S of El Cóndor; 1 Sep. 1981; D.K. Faulkner leg.; SDNHM • 1 ♀; 4 mi S of El Cóndor; 28 May 1981; Faulkner and Brown leg.; SDNHM • 1 ♀; 8 mi E of Tecate on Hwy 2; 30 Jun. 1980; Faulkner and Brown leg.; SDNHM • 1 ♀; 19 mi E of Ojos Negros; 21 Jul. 1980; Faulkner and Brown leg.; SDNHM • 9 ♀♀; Agua Caliente (San Carlos), 18.5 km E of Maneadero; 6 Jul. 1973; P.H. Arnaud, Jr. leg.; CAS • 2 ♀♀; El Pabellón; 2 Jul. 1968; J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU • 1 ♀; Los Mártires; 24 Jun. 1968; N. Leppla, J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU • 1 ♀; Río San Salvador at Hwy 3; 31.8733° N, 116.0908° W; 7–12 Jun. 1997; M.E. Irwin and E.I. Schlinger leg.; INHS 387491 • 19 ♀♀; San Ángel; 28 Jun. 1968; N. Leppla, J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU • 1 ♀; same data as for preceding; 28 Jun. 1968; N. Leppla, J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU M.G.R. Database No. 1215 • 1 ♀; Santa Ana; 21 Jun. 1968; N. Leppla, J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU • 1 ♀; Sierra de Juárez (6 mi N of Laguna Hanson); 21 Jul. 1980; Faulkner and Brown leg.; SDNHM. – Baja California Sur • 1 ♀; 2 mi S of La Paz; 2 Aug. 1966; J.A. Chemsak leg.; EMEC • 1 ♀; 11 mi N of San Isidro (La Purísima); 3 Apr. 1985; Bloomfield and Faulkner leg.; SDNHM • 1 ♀; 36 km NE of San José del Cabo; 8 Mar. 1988; W.J. Pulawski leg.; CAS • 1 ♂; Arroyo San Bartolo; 28 Aug. 1982; D.K.F. leg.; SDNHM • 1 ♀; Loreto 21 Jun. 1983; R. Miller and L. Stange leg.; FSCA • 1 ♀; Playa Los Cerritos (11.2 mi S of Todos Santos); 28 Sep. 1981; F. Andrews and D. Faulkner leg.; SDNHM • 3 ♀♀; Puerto Escondido; 15 Jun. 1992; R. Shaver leg.; UCBME • 1 ♀; S of El Rosario on Hwy 1 at km 115 marker; 24–25 Aug. 1992; D.E. Russell leg.; UCBME • 1 ♀; San Ignacio; 26 Jun. 1938; Michelbacher and Ross leg.; CAS • 2 ♀♀; San José Viejo; 17 Jul. 1968; J. Davidson, J. Bigelow, M. Bentzien, W. Fox, S. Williams, and M. Cazier leg.; ASU. – Chihuahua • 1 ♀; 10–15 km S of Maycoba; 29 Aug. 1984; W.J. Pulawski leg.; CAS. – Coahuila • 2 ♀♀; 61 mi N of Saltillo; 11 Aug. 1959; A.S. Menke and L.A. Stange leg.; UCBME • 1 ♀; Guadalupe; 24 May 1952; M. Cazier, W. Gertsch, and R. Schrammel leg.; AMNH • 1 ♀; Rancho Las Lilas (Reserva de la Biosfera Mapimí); 26.7372° N, 103.7542° W; 12 Jun. 1995; L. Godínez leg.; UNAM LG-1095 7400. – Durango • 1 ♂; Navajos (20 mi E of El Salto); 23 Jul. 1964; L.A. Kelton leg.; CNC 808392. – Guanajuato • 2 ♀♀; León; 16 Aug. 1953; C. and P. Vaurie leg.; AMNH. – Jalisco • 1 ♀; Guadalajara; Crawford leg.; USNM. – Michoacán • 1 ♀; Cotija; 6 Sep. 1970; B.L. Villegas leg.; UCBME. – Nuevo León • 2 ♀♀; 50 mi SE of Monterrey; 13 Oct. 1957; H.A. Scullen leg.; OSAC. – Sonora • 1 ♀; Bahia San Pedro, Gulf of California; 7 Jul. 1921; E.P. Van Duzee leg.; CAS. USA – Arizona • 1 ♀; 0.5 mi W of Hwy 92 on Ash Canyon Road (Huachuca Mountains), Cochise County; 13–22 Sep. 1993; N. McFarland leg.; CNC 1801913 • 1 ♀; 1 mi S of Portal (Chiricahua Mountains), Cochise County; 26 Aug. 1966; M.A. Cazier leg.; ASU • 1 ♀; 11 mi SW of Eloy, Pinal County; 19 Jun. 1953; T.R. Haig leg.; UCBME • 1 ♀; 12 km NE of Tombstone, Cochise County; 31.8460° N, 110.0030° W; 7 Sep. 2009; C.S. Sheffield and S. Dumesh leg.; PCYU PCYU-SD09:1064 • 1 ♀; 12 mi N of Yuma; 8 Oct. 1958; T.R. Haig leg