Paraskusella Cranston, 2018 and Kribiodosis Kieffer, 1921 (Diptera: Chironomidae: Chironomini): two new species from Borneo described, allowing incorporation into a multi-gene molecular phylogeny

Based on Chironomidae (Diptera) specimens collected from Gunung Mulu National Park, Sarawak, East Malaysia, an additional two genera are recognised, belonging to the cosmopolitan tribe Chironomini. Two new species, each represented by adult males, are described and illustrated as Paraskusella sarawak, new species and Kribiodosis mulu, new species. The larva of Kribiodosis Kieffer, 1921 is described for the first time based on its barcode identity to a male of Kribiodosis cantonensis Tang, 2021. New insights into phylogenetic relationships are provided for Paraskusella Cranston, 2018 and Kribiodosis Kieffer, 1921 based on multi-gene analyses that include these taxa. Each is located independently within a broad informal ‘Microtendipes–group’. Biogeographic implications of these ‘tropical Gondwanan’ distributions are discussed briefly.