Cephennodes asymmetricus Jałoszyński, 2025, sp. nov.

Cephennodes (s. str.) asymmetricus sp. nov. (Figs 73–80) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels:“CHINA: Yunnan, SE Pingbian, / 22º54’31’’N, 103º41’44’’E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28.VIII.2014, / leg. M. Schülke [CH14-22 a]” [white, printed]; “ CEPHENNODES (s. str.) / asymmetricus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (37 exx.): 6 ♂♂, 23 ♀♀, same data as for holotype; 4 ♂♂, 4 ♀♀, “CHINA: Yunnan, SE Pingbian, / 22º54’31’’N, 103º41’44’’E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 27.VIII.2014, / leg. M. Schülke [CH14-22]” [white, printed]; each with yellow “ PARATYPUS ” label (MNB, cPJ). Diagnosis. Body large, BL> 1.6 mm; antennal club pentamerous; punctures on frontal region of head and on pronotum larger and deeper than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.25 × EL. Male: antennomere 7 (Figs 74–75) longer and wider than 8, asymmetrical, with flat round mesal region delimited by sharp ridge; frontal region of head between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76); aedeagus (Figs 77–80) in ventral view with apex of capsular region broadly subtriangular and rounded, distal plate elongate, subtriangular and strongly curved dorsad. Female: antennomere 7 distinctly enlarged but symmetrical, longer and slightly broader than 8. Description. Body of male (Fig. 73) strongly convex, oval, with barely marked constriction between pronotum and elytra, light brown with slightly lighter distal regions of legs, covered with yellowish setae. BL 1.68–1.83 mm. Head (Fig. 76) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.25–0.28 mm, HW 0.43–0.45 mm; frontovertexal region weakly convex; frons weakly convex, between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76, indicated by arrowhead); supraantennal tubercles weakly elevated. Punctures on area anterior to eyes distinct and sharply marked, unevenly distributed, on clypeus sparser, close to antennal fossae denser; setae on frons and vertex sparse, long and suberect. Antennae (Figs 73–75) long and slender, with pentamerous clubs, AnL 0.95–1.00 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3–6 each about as long as broad, 7 enlarged and asymmetrical, longer and broader than both 6 and 8, with flat round area on mesal side delimited by sharp ridge, 8 weakly transverse, 9 and 10 each indistinctly transverse, 11 nearly as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest just anterior to base; PL 0.50–0.55 mm, PW 0.75–0.83 mm.Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded and barely noticeably microserrate, posterior corners indistinctly obtuse-angled; posterior margin shallowly bisinuate; lateral marginal carinae narrow and mesally not accentuated by groove, simple (i.e., not doubled). Punctures on median region of pronotal disc large and deep, with sharp margins, separated by spaces subequal to diameters of punctures; punctures reducing in size and depth toward anterior and posterior pronotal margins, those in anterior corners indistinct, in posterior corners denser than those at middle. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.93–1.03 mm, EW 0.80–0.83 mm, EI 1.15–1.28; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.25 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located at middle between lateral margin of mesoscutellum and subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae distinctly longer than those on pronotum, dense and erect. Hind wings lacking. Metaventrite lacking lateral impressions, convex. Legs long and slender, unmodified. Aedeagus (Figs 77–80) of modified simonis form, AeL 0.48 mm; capsular region of median lobe in ventral view approximately drop-shaped, stout, with subtriangular and broadly rounded apex shifted to the left side; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at weakly obtuse angle; parameres long and slender, not reaching apex of aedeagus, each with three long subapical setae. Female. Externally similar to male, except for antennomere 7 distinctly enlarged but symmetrical, head dorsum unmodified, and slightly shorter antennae in relation to BL; wingless; BL 1.65–1.78 mm; HL 0.25 mm, HW 0.53– 0.58 mm, AnL 0.90–0.93 mm; PL 0.55–0.58 mm, PW 0.78–0.85 mm; EL 0.88–0.98 mm, EW 0.78–0.83 mm, EI 1.09–1.19. Distribution. S China: Yunnan. Etymology. The adjective asymmetricus refers to the asymmetrical antennomere 7 in males. Remarks. Only three species of the C. impressifrons group have the antennomere 7 modified in males, and only two of them have this antennomere bearing a round mesal flat area: C. antennatus and C. asymmetricus (the third species, C. triangularis described below, has a subtriangular mesal process on antennomere 7). The Chinese Cephennodes asymmetricus and Vietnamese C. antennatus have clearly different modifications of the head in males. In the latter species (illustrated in Jałoszyński & Nomura (2009): fig. 110), there is a pair of minute frontal tubercles at middle of a large virtually impunctate area delimited anteriorly from a strongly punctate clypeofrontal region by an arcuate transverse ridge, whereas in C. asymmetricus there is neither a ridge nor tubercles (Fig. 76). Moreover, in C. antennatus, the apex of the capsular region of the median lobe forms a strongly elongate and slender rod-like process, whereas in C. asymmetricus the apex is broadly rounded; the distal plate in C. antennatus is broadening distad, vs. tapering in C. asymmetricus.

Publication Details

Journal
Zenodo (CERN European Organization for Nuclear Research)
Published
2025-07-18
DOI
https://doi.org/10.5281/zenodo.16606480
Primary Topic
Subterranean biodiversity and taxonomy
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article

Cephennodes asymmetricus Jałoszyński, 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Subterranean biodiversity and taxonomy
article

Cephennodes asymmetricus Jałoszyński, 2025, sp. nov.

article en

Abstract

Cephennodes (s. str.) asymmetricus sp. nov. (Figs 73–80) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels:“CHINA: Yunnan, SE Pingbian, / 22º54’31’’N, 103º41’44’’E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28.VIII.2014, / leg. M. Schülke [CH14-22 a]” [white, printed]; “ CEPHENNODES (s. str.) / asymmetricus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (37 exx.): 6 ♂♂, 23 ♀♀, same data as for holotype; 4 ♂♂, 4 ♀♀, “CHINA: Yunnan, SE Pingbian, / 22º54’31’’N, 103º41’44’’E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 27.VIII.2014, / leg. M. Schülke [CH14-22]” [white, printed]; each with yellow “ PARATYPUS ” label (MNB, cPJ). Diagnosis. Body large, BL> 1.6 mm; antennal club pentamerous; punctures on frontal region of head and on pronotum larger and deeper than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.25 × EL. Male: antennomere 7 (Figs 74–75) longer and wider than 8, asymmetrical, with flat round mesal region delimited by sharp ridge; frontal region of head between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76); aedeagus (Figs 77–80) in ventral view with apex of capsular region broadly subtriangular and rounded, distal plate elongate, subtriangular and strongly curved dorsad. Female: antennomere 7 distinctly enlarged but symmetrical, longer and slightly broader than 8. Description. Body of male (Fig. 73) strongly convex, oval, with barely marked constriction between pronotum and elytra, light brown with slightly lighter distal regions of legs, covered with yellowish setae. BL 1.68–1.83 mm. Head (Fig. 76) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.25–0.28 mm, HW 0.43–0.45 mm; frontovertexal region weakly convex; frons weakly convex, between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76, indicated by arrowhead); supraantennal tubercles weakly elevated. Punctures on area anterior to eyes distinct and sharply marked, unevenly distributed, on clypeus sparser, close to antennal fossae denser; setae on frons and vertex sparse, long and suberect. Antennae (Figs 73–75) long and slender, with pentamerous clubs, AnL 0.95–1.00 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3–6 each about as long as broad, 7 enlarged and asymmetrical, longer and broader than both 6 and 8, with flat round area on mesal side delimited by sharp ridge, 8 weakly transverse, 9 and 10 each indistinctly transverse, 11 nearly as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest just anterior to base; PL 0.50–0.55 mm, PW 0.75–0.83 mm.Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded and barely noticeably microserrate, posterior corners indistinctly obtuse-angled; posterior margin shallowly bisinuate; lateral marginal carinae narrow and mesally not accentuated by groove, simple (i.e., not doubled). Punctures on median region of pronotal disc large and deep, with sharp margins, separated by spaces subequal to diameters of punctures; punctures reducing in size and depth toward anterior and posterior pronotal margins, those in anterior corners indistinct, in posterior corners denser than those at middle. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.93–1.03 mm, EW 0.80–0.83 mm, EI 1.15–1.28; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.25 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located at middle between lateral margin of mesoscutellum and subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae distinctly longer than those on pronotum, dense and erect. Hind wings lacking. Metaventrite lacking lateral impressions, convex. Legs long and slender, unmodified. Aedeagus (Figs 77–80) of modified simonis form, AeL 0.48 mm; capsular region of median lobe in ventral view approximately drop-shaped, stout, with subtriangular and broadly rounded apex shifted to the left side; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at weakly obtuse angle; parameres long and slender, not reaching apex of aedeagus, each with three long subapical setae. Female. Externally similar to male, except for antennomere 7 distinctly enlarged but symmetrical, head dorsum unmodified, and slightly shorter antennae in relation to BL; wingless; BL 1.65–1.78 mm; HL 0.25 mm, HW 0.53– 0.58 mm, AnL 0.90–0.93 mm; PL 0.55–0.58 mm, PW 0.78–0.85 mm; EL 0.88–0.98 mm, EW 0.78–0.83 mm, EI 1.09–1.19. Distribution. S China: Yunnan. Etymology. The adjective asymmetricus refers to the asymmetrical antennomere 7 in males. Remarks. Only three species of the C. impressifrons group have the antennomere 7 modified in males, and only two of them have this antennomere bearing a round mesal flat area: C. antennatus and C. asymmetricus (the third species, C. triangularis described below, has a subtriangular mesal process on antennomere 7). The Chinese Cephennodes asymmetricus and Vietnamese C. antennatus have clearly different modifications of the head in males. In the latter species (illustrated in Jałoszyński & Nomura (2009): fig. 110), there is a pair of minute frontal tubercles at middle of a large virtually impunctate area delimited anteriorly from a strongly punctate clypeofrontal region by an arcuate transverse ridge, whereas in C. asymmetricus there is neither a ridge nor tubercles (Fig. 76). Moreover, in C. antennatus, the apex of the capsular region of the median lobe forms a strongly elongate and slender rod-like process, whereas in C. asymmetricus the apex is broadly rounded; the distal plate in C. antennatus is broadening distad, vs. tapering in C. asymmetricus.

Zenodo (CERN European Organization for Nuclear Research)
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Subterranean biodiversity and taxonomy
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