Medioptyx Volynkin & Černý, 2024, gen. n.

Genus Medioptyx gen. n. https://zoobank.org/ urn:lsid:zoobank.org:act: A28520EA-CDF5-49E7-8CDC-6628CC22805C Type species: Medioptyx kishidai sp. n., by present designation. Diagnosis. The only known species of the new genus (Figs 25, 26) has forewing shape, pattern and venation very similar to Collitoptyx (Figs 1–24) but the significant differences in both male and female genitalia prove its generic status. The male genitalia of Medioptyx gen. n. (Figs 46, 47) are characterised by the combination of the following features. (1) The conjuga is triangular proximally, rod-like distally, articulated with its counterpart, and both the processus momenti and the tendon are equally well-sclerotised (similar to Caliginoptyx gen. n.) whereas in Collitoptyx the tendon part performs a skeletal function. (2) The anellus is thick-walled and bears two broad ventro-lateral plates (similar to Collitoptyx (Insolitoptyx) subgen. n.). (3) The main diverticulum of the vesica bears a sclerotised plate armed with several spike-like cornuti (vs. a single conical cornutus in Collitoptyx). (4) The sub-ejaculatorial (apical) section of the vesica is proximally enlarged, bears a thorn-like cornutus, and the vesica ejaculatorius originates from its distal part (similar to Thysanoptyx) whereas the vesica ejaculatorius of Collitoptyx originates from the base of the vesica. The female genitalia of Medioptyx gen. n. (Fig. 61) differ from Collitoptyx (Figs 52–60) in the long, dorso-ventrally flattened and heavily sclerotised ductus bursae (similar to Thysanoptyx), which is short or reduced and entirely or posteriorly membranous in Collitoptyx. Compared to Thysanoptyx (Fig. 62), the female genitalia of the new genus have a gelatinous and rugose posterior section of the corpus bursae with a broad ventral protrusion bearing a posterior sclerotised area, and a membranous and more or less globular anterior section, which is similar to Thysanoplisna but in the latter it also bears a signum, which is absent in the new genus. Etymology. The genus name is an aggregate of the Latin adjective ‘medius’ meaning ‘intermediate’ and the genus- group name Thysanoptyx. The name refers to the genitalia morphology of the new genus, which combines the features characteristic of Collitoptyx and Thysanoptyx. Gender is feminine. Species content. The genus is monotypic.