Hypselostoma khaowongense Panha 1998

Hypselostoma khaowongense Panha, 1998 Hypselostoma khaowongensis Panha, 1998 b: 66, fig. 4. Hypselostoma khaowongensis — Hemmen and Hemmen 2001: 41. Gyliotrachela saraburiensis Panha & Burch in Burch et al. 2003: 143–151, figs 6–7. syn. nov. Gyliotrachela muangon Panha & Burch in Panha et al. 2004: 67–68, fig. 7. syn. nov. Gyliotrachela khaowongensis — Panha and Burch 2008: 71–72, fig. 62; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8. Gyliotrachela saraburiensis — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 78–80, fig. 68; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8. Gyliotrachela muangon — Panha and Burch 2008: 76–77, fig. 66; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8; Gojšina et al. 2022: 137, fig. 6; Tongkerd et al. 2024: 182–184, figs 11,13O. Type material examined. Thailand • holotype; S. Panha leg.; CUMZ ver. 001 • paratype of G. saraburiensis; from the type locality; CUMZ 44047 • 2 paratypes of H. khaowongense; from the type locality; SMF 331439 • 3 paratypes of G. saraburiensis; from the type locality; SMF 331446 • 2 paratypes of G. muangon; from the type locality; S. Panha leg.; SMF 331459. Additional material examined. Thailand-North • 24 shells; Chiang Rai Province, Chiang Rai ca 5 km west from centre of Wat Phra That Tham Doi Kong Khao; 9°54.820'N, 99°46.642'E; 11 Feb. 2015; A. Hunyadi leg.; coll. HA • 33 shells; Chiang Rai Province, ca 5 km WNW of Chiang Rai, Tham Phra cave by Mae Nam Kok river, limestone rocks near the cave entrance; 19°55.047'N, 99°47.342'E; 420 m a. s. l; Sep. 2007; A. Reischütz leg.; coll. REI • 12 shells; Chiang Mai Province, 5 km northwest from Chiang Dao, vicinity of Wat Tham Chiang Dao; 19°23.628'N, 98°55.711'E; 450 m a. s. l.; 07 Feb. 2015; A. Hunyadi leg.; coll. HA • 13 shells; Chiang Mai Province, Doi Pha Sao Mountain, 3 km W of Ban Prang Mao; 19°25.98'N, 99°3.48'E; 500 m a. s. l.; 19 Jun. 1987; F. G. Thompson leg.; locality code FGT-4338; UF 528706 • 21 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with Dipterocarp forest, leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20 May 1988; F. G. Thompson leg.; locality code FGT-4453; UF 347006 • 59 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with Dipterocarp forest, leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20. May 1988; F. G. Thompson leg.; locality code FGT-4452; UF 346992 • 150 shells; Mae Hong Son Province, 13.9 km NW Pai, along road 1095; 19°23'N, 98°23'E; 780 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0571; UF 345938 • 3 shells; Mae Hong Son Province, Ban Tham Lod Village, 7 km N Soppong on Road 1095; 19°34'N, 98°9'E; 790 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0574; UF 591328 • 20 shells; Mae Hong Son Province, 21 km N Mae Hong Son, Fish Cave; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0593; UF 345672 • 40 shells; Mae Hong Son Province, 40 km N of Mae Hong Son; 19°27'N, 97°59'E; 550 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0588; UF 345643 • 1 shell; Mae Hong Son Province, 21 km N Mae Hong Son, Fish Cave; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 A; UF 345702 • 7 shells; Mae Hong Son Province, Mae Hong Son, Road 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 B; UF 591329 • 3 shells; Mae Hong Son Province, Mae Hong Son, Road 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595; UF 345699 • 1 shell; Lampang Province, 6 km SW Mae Pak, limestone hill; 17°57.8833'N, 98°51.1167'E; 600 m a. s. l.; 13 June 1987; F. G. Thompson leg.; locality code FGT-4328; UF 591330 • 11 shells; Lampang Province, limestone dome at pass 11 km SSW of Ban Pang La; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.; 14 May 1988; F. G. Thompson leg.; locality code FGT-4441; UF 591331 • 2 ethanol preserved shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4440; UF 346908 • 43 shells; Phrae Province, limestone knoll 4 km N of Ban Nam Rin, deep rocks and leaves; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4444; UF 346931 • 1 shell; Phrae Province, Than Pha Nan Cave, 6 km NE of Rang Kwang; 18°23.1'N, 100°20.7667'E; 300 m a. s. l.; 30 Apr. 1988; F. G. Thompson leg.; locality code FGT-4400; UF 347403 • 2 shells; Phrae Province., 4 km N of Ban Nam Rin valley with limestone ridges, leaf litter at base of boulders; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4445; UF 346935 • 5 ethanol preserved shells; Phrae Province, 11 km W of Phrae, on Road 1023 teak plantation and cut forest; 18°10.6833'N, 100°4.2333'E; 290 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4446; UF 346967 • 19 shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4447; UF 591332. Thailand-Northeast • 28 shells; Petchabun Province, 24.4 km S. Petchabun on Hwy. 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0693; UF 346096 • 1 shell; Nakhon Ratchasima Province, 3.4 km W of Ban Mu Si; 14.534°N, 101.380°E; 380 m a. s. l.; 05. May 1987; F. G. Thompson leg.; locality code FGT-4241; UF 534868 • 9 shells; Nakhon Ratchasima Province, Pak Chong district, mountain above Wat Thep Phithak Punnaram; 14°36.865'N, 101°15.926'E; 500 m a. s. l.; 04 Mar. 2023; A. Hunyadi leg.; coll. HA. Thailand-West • 16 shells; Kanchanaburi Province, Erawan Nat. Park, Erawan Falls Trail; 14°22.310'N, 99°08.699'E; 85 m a. s. l.; 17 Feb. 2015; A. Hunyadi leg.; coll. HA • 10 shells; Petchabun Province, 24.4 km S. Petchabun on Hwy. 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0694; UF 346100. Thailand-Central • 10 shells; Phitsanulok Province, Noen Maprang district, Chomphu, Tham Phra Wang Daeng; 16°40.651'N, 100°41.287'E; 02 Mar. 2023; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 37 shells; Nakhon Sawan Province, 19.5 km NE intersection of Hwys 11 and 32, limestone outcrop, base of cliff under Georissa zone; 15°14'N, 100°16'E (provided coordinates point in Chai Nat Province); 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697; UF 346135 • 1 shell; Nakhon Sawan Province, 19.5 km NE intersection of Hwys 11 and 32; 16°14'N, 100°16'E; 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697; UF 346126 • 1 shell; Lopburi Province, Tha Wung district, Khao Samo Khon, Wat Tham Chang Pueak; 14°54.139'N, 100°30.151'E; 30 m a. s. l.; 03 Mar. 2023; A. Hunyadi leg.; coll. HA • 2 shells; Saraburi Province, behind Phra Phutthabat, limestone hill with scrubby evergreen forest, leaf litter among rocks; 14.733°N, 100.800°E; 50 m a. s. l.; 03. May 1988; K. Auffenberg leg.; locality code KA-0699; UF 508979. Laos • 27 shells; Luang Prabang Province, old forest on Phousi Mountain in Luang Prabang, clay, black soil, on and under rocks; 19°53.411'N, 102°08.216'E; 374 m a. s. l.; 13 Nov. 2004; A. Abdou & I. Muratov leg.; MNHN -IM-2012-25387 • 6 shells; Luang Prabang Province, 3.1 km from centre of Nong Khiaw towards Pak Xeng, Tham Pha Toke, vicinity of the cave; 20°33.215'N, 102°37.722'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 24 shells; Luang Prabang Province, 12 km from centre of Nong Khiaw towards Pak Xeng, northeast from Ban Huai Lek, left side of the road; 20°32.585'N, 102°41.161'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA. Myanmar • 3 shells; Shan State, c. 3.6 km NNE of Hsihseng centre, limestone hill; 20°11′15″N, 97°16′07″E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 7 shells; Shan State, 9.5 km from centre of Hsihseng; 20°14.066'N, 97°15.514'E; 1140 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 21 shells; Shan State, 3.6 km north-northeast from centre of Hsihseng; 20°11.248'N, 97°16.114'E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 63 shells; Shan State, ca 16 km from centre of Taunggyi centre towards Hopong, 1.5 km north on rd. # 4, „ Shwe Pyi Aunchonda ” monastery; 20°47.263'N, 97°8.239'E; 1110 m a. s. l.; 08 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA • 57 shells; Shan State, ca 6 km east from centre of Hsihseng; 20°8.002'N, 97°18.024'E; 500 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA. Type localities. “ A limestone hill near Takli District, Nakornsawan Province, 15°16′52″N, 100°22′38″E, 114 meters elevation ”, Thailand (H. khaowongense); “ East of Tepitak Tamaram Temple, Tepitak Mountain, Muang Saraburi District, Saraburi Province, 14°36'57"N, 101°15'50"E, 510 meters elevation. Thailand ” (G. saraburiensis); “ Muangon Cave, San Kam Pang District, Chiangmai Province, 18°47'11"N, 99°14'16"E, 280 meters elevation ” (G. muangon). Differential diagnosis. This species is similar to H. holimanae but is clearly different by the separated angular and parietal lamellae (fused in H. holimanae) as well as the presence of numerous and variable apertural barriers (only four in H. holimanae). Hypselostoma fruhstorferi has a narrower umbilicus and fewer apertural barriers. This species is also similar to H. cultura from which it can be separated by the less prominently shouldered last whorl and a narrower umbilicus. Distribution. Clearly a widespread species: its distribution stretches from Chiang Rai province in N Thailand to the southernmost known localities in Ratchaburi and Chonburi provinces. Westwards, this species stretches to Taunggyi in Myanmar and the easternmost locality is known from Luang Prabang province in Laos. Remarks. Examined specimens h