Tumeochrysa (Nineta) grandis Wu & Liu 2024, comb. nov.

[29] Tumeochrysa (Nineta) grandis (Navás, 1915) comb. nov. (Figs 36, 84) Nineta grandis Navás, 1915a: 395. Type locality: China (Manchuria, Región del Lago Hanka [Heilongjiang, Xingkai Lake]). Holotype in MNHN. Diagnosis. This species is characterized by the presence of the head and thorax without spots, the outer gradate series of forewing darker than inner gradates series, the male sternum IX slightly elongated and curved dorsal apically, and the female subgenitale rounded and bilobed apically. Description. Male. Body length 19.0– 20.9 mm, forewing length 19.3–22.5 mm, hindwing length 17.1–19.7 mm. Head without spots, vertex raised. Compound eyes prominent and black. Antennae yellow, scape and pedicel wider than flagellum. Frons, clypeus and gena without spots. Maxillary palpi with brown stripe on outer side, labial palpi brownish yellow. Pronotum anterior margin shorter than posterior margin, with pale brown transverse markings. Mesonotum and metanotum without markings. Legs with dense setae and without spots. Pretarsal claws brown, with basal dilation. Forewing broad, without shadings. Pterostigma indistinct. Costal area with 27–32 crossveins; 18–19 radial crossveins present. m1 smaller than m2; im narrow and ovoid; 10–11 psm-psc crossveins present. c1 smaller than c2, dcc opened. A1 forked, A2 and A3 simple. CuP brown. Inner gradate series with 10–13 crossveins, outer gradate series with 12 brown crossveins. Hindwing narrow. Pterostigma indistinct. Costal area with 23–26 crossveins; 17–20 radial crossveins and 10–11 psm-psc crossveins present. Inner gradate series with 9–11 crossveins, outer gradate series with 11–13 crossveins. Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII and sternum VIII about 1/3 as long as tergum VII; sternum IX slightly elongated and curved towards dorsal apically; tergum IX fused with ectoproct; ectoproct rounded. Callus cerci ovoid. Gonarcus arcuate, median arch with broad U-shaped invagination, lateral arms flat and expanded; entoprocessus broad; arcessus rounded at base, pointed at tip. Female. Body length 16.1–19.0 mm, forewing length 19.7–23.9 mm, hindwing length 17.6–22.5 mm. Sternum VII slightly longer than tergum VII; tergum VIII obviously shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically slightly; spermatheca thin; vela short; ventral impression absent; duct long and coiled. Material. 6♀, China, Beijing, Miyun, Yunmengshan National Forestry Park, light, 2013.IX.09, Yuqiang Xi (CAU); 2♂ 1♀, China, Beijing, Mentougou District, Baihuashan National Nature Reserve, 1098 m, 2022.VI.15–17, Xingyue Liu (CAU); 2♂ 12♀, China, Beijing, Yanqing District, Songshan National Nature Reserve, 2012.IX.07–08, Xiao Zhang (CAU); 6♀, China, Beijing, Yanqing District, Songshan National Nature Reserve, 2012.IX.07, Yuqiang Xi (CAU); 2♀, China, Hebei, Chengde, Mulan Paddock, light, 1180 m, 2016.VII.10 (CAU); 1♀, China, Hebei, Zhangjiakou, Heilongshan Forestry Station, light, 1331 m, 2016.VII.05 (CAU); 2♂ 2♀, China, Hebei, Chengde, Wulingshan National Nature Reserve, 2021.VIII.20–21, Yan Lai (CAU); 1♂ 15♀, China, Neimenggu, Hulunbeier, Hanma National Nature Reserve, 2017.VIII (CAU); 1♂ 1♀, China, Neimenggu, Hulunbeier, Lanqimiao District, 2017.VII.17, Ya’nan Lv (CAU); 2♀, China, Neimenggu, Chifeng, Sanhanwula National Nature Reserve, 1200 m, 2013.VII.24, Xiumei Lu (CAU); 1♀, China, Liaoning, Benxi, Laotudingzi National Nature Reserve, light, 570 m, 2014.VIII.20, Tong Zhang (CAU); 1♀, China, Liaoning, Benxi, Tanggou Scenic Area, 600 m, 2022.VIII.01, Xingyue Liu (CAU); 2♂ 1♀, China, Heilongjiang, Tongjiang, Hezhezu, 110 m, 2022.VII.17, Xueyuan Wang (CAU); 4♀, China, Heilongjiang, Tongjiang, Hezhezu, Jiangshengtun, 116 m, 2022.VII.18, Xueyuan Wang (CAU); 7♂ 15♀, China, Heilongjiang, Heihe, Wudalianchi Scenic Area, 344 m, 2022.VII.23 (CAU); 8♂ 4♀, China, Heilongjiang, Yichun, Liangshui National Nature Reserve, 320–400 m, 2022.VII.21–22, Yuchen Zheng (CAU); 3♂ 2♀, China, Heilongjiang, Yichun, Fenglin, Wuying, Fenglin National Nature Reserve, 288 m, 2022.VII.19, Xueyuan Wang (CAU); 2♂ 3♀, China, Heilongjiang, Fuyuan, Wusu, 2022.VII.15–16, Xueyuan Wang (CAU); 2♂, China, Heilongjiang, Mishan, Dangbi, Xingkai Lake, 2022.VII.14–15, Yuchen Zheng (CAU); 1♀, China, Heilongjiang, Muling, Gonghe, Mt. Liufengshan, 450 m, 2022.VII.11, Yuchen Zheng (CAU); 1♀, China, Heilongjiang, Nenjiang, Sichilihe Forestry Station, 2014.VII.21, Ya’nan Lv (CAU); 1♀, China, Heilongjiang, Nenjiang, Qianjin, 2014. VII.22, Ziqiang Sun (CAU); 1♂ 1♀, China, Heilongjiang, Heihe, Wudalianchi Scenic Area, 1985.VIII.07–08, Chikun Yang (IOZ); 1♀, China, Heilongjiang, Ning’an, Jingpohu National Scenic Area, 1985.VIII.15, Chikun Yang (IOZ); 1♀, China, Sichuan, Jiangyou, Wanglang National Nature Reserve, 2596 m, 2015.VII.06, Lu Yue (CAU); 2♀, China, Shaanxi, Xi’an, Zhouzhi, Houzhenzi, light, 2018.VII.14–15, Yingnan He (CAU); 5♀, China, Shaanxi, Baoji, Taibaishan National Nature Reserve, 1126 m, 2018.VIII.24, Yuqiang Xi (CAU); 1♀, South Korea, Kangwon-do, Tacbeak, Mt. GeomRyongSo, light, 768 m, 2013.IX.12, Xuankun Li (CAU); 2♀, South Korea, Kangwon-do, Yangyang, Mt. Jeombong, light, 495 m, 2013.IX.11, Xuankun Li (CAU); 3♀, South Korea, Kangwon-do, Mt. Inje Daream, 574 m, 2013.IX.10, Xuankun Li (CAU). Distribution. China (Beijing, Hebei, Heilongjiang, Liaoning, Neimenggu, Shaanxi, Sichuan); South Korea (Kangwon-do). Remarks. This species is similar to T. breva sp. nov. and T. shaanxiensis. The invagination on the median arch of male gonarcus of this species is trapezoid, whereas in the latter two it is U-shaped. In addition, the female subgenitale of this species is more rounded and slightly bilobed. The female subgenitale of T. shaanxiensis and T. breva sp. nov. is heart-shaped and more squared, respectively.