Trichastylopsis skillmani Santos-Silva, Perez-Flores, and Botero. 2024, New species

Trichastylopsis skillmani Santos-Silva, Pérez-Flores, and Botero. New species (Fig. 4) ZooBank Registration number: urn:lsid:zoobank.org:act: BD4E63C9-EF9A-47DC-8977- F4775C33644F Description. Holotype female: Integument mostly dark brown, almost black on some areas; ventral mouthparts light brown, except brown mentum and palpomeres dark brown with light brown apex; anteclypeus brownish; labrum light brown on posterior third, yellowish brown on anterior 2/3; antennomere 3 with irregular brown areas; antennomere 4 with irregular brown areas on basal 2/3 and dark reddish-brown ring on apical third; antennomeres 5–10 reddish brown except dark brown dorsal macula on base, reaching sides on some antennomeres, and dark brown apex, light area gradually shorter towards antennomere 10 (missing antennomere 11). Apex of ventrites 1–4 brownish. Head. Frons densely, finely punctate; with dense pale beige pubescence close to clypeus and eyes, gradually light brown with yellowish-white interspersed on remaining surface, except white pubescent macula centrally close to vertex; with one long, erect seta close to eyes, setae black basally, yellowish white on remaining surface; with sparse, short, erect yellowish-white setae interspersed. Area between antennal tubercles and eyes with dense yellow pubescence. Vertex with dense, both brown and yellowish-brown pubescence partially obscuring integument, except glabrous median groove and yellow pubescence close to eyes. Area behind upper eye lobes with abundant pubescence almost obscuring integument on some areas, pubescence pale yellow, except brownish pubescent macula after middle; with a few long, erect setae close to eye, setae black basally, yellowish white on remaining surface. Area behind lower eye lobes with dense pale-yellow pubescence, except glabrous area about middle. Genae 1.3 times longer than lower eye lobe; with abundant yellowish-white pubescence not obscuring integument, except dense pale-yellow pubescence close to eye, sparser yellowish-brown pubescence on centre of frontal region, and glabrous apex; with sparse, long, erect yellowish-white setae interspersed, some setae black basally. Antennal tubercles moderately elevated; with abundant pubescence partially obscuring integument, pubescence mostly beige frontally, mostly yellowish brown on remaining surface. Wide central area of postclypeus with abundant beige pubescence partially obscuring integument close to frons, sparser close to anteclypeus; with a few long, erect beige setae interspersed close to frons, and moderately abundant, long setae directed forwards close to anteclypeus, setae mostly yellowish white centrally, pale yellow laterally. Sides of postclypeus glabrous. Labrum with abundant white pubescence not obscuring integument on posterior third, glabrous on anterior third, except fringe of yellow setae on anterior margin; with long, erect pale-yellow setae directed forwards posteriorly close glabrous area, some setae black basally. Outer side of mandibles with dense pale-yellow pubescence on posterior third, with a few long, erect setae interspersed, setae black basally, pale yellow on remaining surface; remaining surface glabrous. Gulamentum smooth, glabrous, except fine, arched striae anterocentrally, and greyish-white pubescence not obscuring integument on intermaxillary process. Distance between upper eye lobes 0.21 times the distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.52 times the distance between outer margins of eyes; inner margin of upper eye lobes wide, rounded. Antennae (from base to apex of antennomere 10) 1.7 times the elytral length, reaching elytral apex at basal quarter of antennomere 8. Scape with dense yellowish-white pubescence centrally partially obscuring integument, pubescence whiter depending on light intensity, and abundant light brown pubescence not obscuring on basal and apical fifths, except apex of apical fifth with greyish-white pubescence, yellower on outer apex; with long, erect yellowish-white setae ventrally, and a few moderately long, erect yellowish-white setae dorsally. Pedicel mostly with abundant brown pubescence partially obscuring integument; with long, erect pale-yellow setae on posterior half of ventral surface. Antennomere 3 with abundant brown pubescence not obscuring integument and abundant, irregular white pubescent rings interspersed; with long, erect pale-yellow setae ventrally, and sparse, short, erect yellowish-white setae interspersed on remaining surface. Antennomeres 5–7 with dense white pubescence, with irregular light brownish pubescent maculae interspersed, except dark brown pubescent macula on dorsal base and abundant dark brown pubescence not obscuring integument on apical quarter; with sparse, long, erect pale-yellow setae ventrally and sparse, short, erect yellowish-white setae interspersed on remaining surface. Antennomeres 8–10 with dense white pubescence, except brown pubescence not obscuring integument basally, and abundant dark brown pubescence not obscuring integument on apical third; with a few long, erect pale-yellow setae on apex of ventral surface, and sparse, short, erect yellowish-white setae interspersed on remaining surface. Antennal formula (ratio) based on length of antennomere 3: scape = 0.71; pedicel = 0.11; antennomere 4 = 0.74; antennomere 5 = 0.60; antennomere 6 = 0.50; antennomere 7 = 0.44; antennomere 8 = 0.44; antennomere 9 = 0.44; antennomere 10 = 0.40. Thorax. Prothorax wider than long; with large, rounded tubercle about middle. Pronotum with five tubercles, one subconical on each side of anterior third, the most elevated, one small, slightly elevated, rounded, on each side just after middle, another large, subelliptical, slightly elevated, located centrally from about middle to posterior fifth; moderately abundantly and coarsely punctate; sides with dense pale-yellow pubescence except moderately large, dense black pubescent macula on each side close to posterior margin, one small, subrounded black pubescent macula on apex of lateral tubercle of prothorax, one subrounded black pubescent macula on each side near posterior margin of lateral tubercle of prothorax, another very small, rounded black pubescent macula on each side near central area of lateral tubercles of prothorax, one small, irregular brown pubescent macula on each side near posterior margin, and irregular black pubescent macula on each side near anterior margin; central region with abundant light brown pubescence not obscuring integument, except dense white pubescence on sides of this region, white pubescent area wider on posterior half and obliquely, somewhat fragmented projected towards centre, irregular yellowish-brown pubescent macula on each side of anterior quarter, and abundant white pubescence on each side close to anterior margin; with a few long, erect dark brown setae on sides of posterior third. Sides of prothorax with dense yellowish-white pubescence, pubescence appearing to be whiter depending on light intensity, except slightly sparser pubescence close to anterior margin, light brown pubescence under lateral tubercles. Prosternum with dense yellowish-white pubescence laterally, and abundant yellowish-grey pubescence not obscuring integument centrally. Prosternal process laterally projected under procoxae about middle; with abundant yellowish-grey pubescence not obscuring integument; narrowest area before and after central projection 0.55 times procoxal width. Ventral surface of mesothorax and metathorax with dense yellowish-white pubescence laterally and abundant white pubescence partially obscuring integument centrally, except wide central area of mesoventrite with yellowish-white pubescence centrally and glabrous laterally. Sides of mesoventral process convergent from base to near apex, then distinctly widened; apex concave; narrowest area 0.73 times mesocoxal width. Scutellum with dense white pubescence centrally, pubescent area gradually widened from base to apex, and abundant yellowish-brown pubescence not obscuring integument laterally. Elytra. Moderately abundantly, coarsely punctate, punctures coarsely on sides of dorsal surface, finer and sparser on posterior third; apex subrounded; with two centrobasal crests, innermost slightly longer, both carina shaped, with dense, short, erect tufts of both black and yellowish-brown setae, posterior tuft of innermost crest longer; remaining dorsal surface with sparse tufts of short, erect setae, some tufts only with black setae, some with both yellowish-brown and black setae; humeral carina slightly marked; dorsal surface with two longitudinal carinae slightly marked, starting on centrobasal crests, ending near elytral apex, fused on their apices; posterior half with three oblique, parallel-sided carinae, anterior carinae from humeral carina to suture, posterior carina less elevated, not reaching suture; with four longitudinal, narrow yellowish-brown pubescent bands, two along longitudinal carina almost reaching elytral base, innermost starting between inner centrobasal crest and suture, distinctly arched towards suture near its base, then following along suture, another starting on humerus apically fused to the bands on carinae; anterior oblique carinae with abundant light brown pubescence; dorsal area between base and second oblique carinae and anterior 4/5 of area between humeral carina and area close to epipleural margin with abundant pubescence partially obscuring integument, forming irregular areas with white, pale-yellow, light brown, and greyish-yellow pubescence; area between second and third oblique carinae with dense yellow pubescence obscuring integument; dorsal area between third oblique carina and apex and apical fifth of lateral area with abundant pale-yellow pubescence with white pubescent maculae interspersed, pale-yellow pubescence denser close to suture; epipleural margin with white pubescence with dark brown pubescent spots in