Clistopyga muisca Palacio & Bordera 2025, sp. nov.

25. Clistopyga muisca Palacio & Bordera, sp. nov. (Figs 37, 38, 52B, Appendix I) Diagnosis. Clistopyga muisca may be distinguished from other species of the C. calixtoi species group by the following combination of characters: face mostly black (Figs 37B, 38E); genal orbits widely interrupted (Fig. 37C, 38B); mesopleuron mostly orange or reddish (Figs 37C, 38B); propodeum widely tinged yellow laterally (Figs 37I, 37B); face with deep and dense setiferous punctures (Figs 37B, 38E); epomia as a small tubercle or short carina at the bottom of the submarginal depression of pronotum (Fig. 37F, arrow); tergite I with lateromedian longitudinal carinae weak, present at anterior 0.2 of tergite; submetapleural carina only present on anterior part (Figs 37I, 38B). Additionally, female has metapleuron and background of propodeum and metasomal tergites striking light orange (Figs 37C, 37E 37G); metasoma with posterolateral well defined white cream marks on tergites I–V(VI) clearly separated from posterior rim (Fig. 37E, 37G); ventral margin of ovipositor sheath basally without denticles. Male has mesosoma extensively marked with red or orange (Figs 38A, 38B); mesoscutum reddish orange (Fig. 38G); metapleuron mostly orange (Fig. 38B); metasoma dark brown, without posterolateral well defined white cream or yellow spots on tergites (Figs 38C, 38H); gena in frontal view straight or slightly convex in profile, without any traces or longitudinal concavity posterior to mandible base (Fig. 38D–E); tergite II 1.38–1.53× as long as posteriorly broad, with dense, wide and relatively shallow setiferous punctures, setae relatively long (Fig. 38F). Description. Female: Body length 7.0– 9.4 mm. Fore wing length 5.2–6.4 mm. Head (Figs 37B–D). In dorsal view, moderately narrowed behind eyes. Gena shiny with uniformly sparse setiferous punctures, in dorsal view slightly rounded, 0.35–0.49× as long as eye, in frontal view slightly convex, moderately constricted below eyes. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 1.05–1.15× its maximum diameter. Distance between hind ocelli 0.95–1.14× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform behind mandible. Face with deep and dense setiferous punctures, distance between punctures less than twice the diameter of punctures. Clypeal suture strongly curved. Clypeus 1.72–2.15× as broad as medially long, distinctly convex in dorsal half, flat to slightly concave in ventral half, with ventral margin slightly concave to straight. Malar space 0.96–1.1× as long as basal mandibular width, with a granulate shallow subocular sulcus. Antenna with 25–28 flagellomeres, first flagellomere 5.0–5.3× as long as wide. Mesosoma (Figs 37A, 37C–D, 37F, 37I). Pronotum smooth and shiny, with fine and scattered setiferous punctures in dorsal posterior part. Epomia as a short carina at the bottom of the submarginal depression of pronotum. Mesoscutum smooth and shiny, with evenly sparse shallow setiferous punctures, denser on anterior part of median lobe setae short; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus moderately deep, reaching 0.4–0.5 the length of mesoscutum. Mesopleuron shiny, with evenly sparse shallow setiferous punctures on anterior and ventral part. Epicnemial carina weak, sinuous, its dorsal end slightly curved towards the back, ending at level of centre of pronotum. Metapleuron smooth and shiny, glabrous, except for some sparse setiferous punctures at posterior part, 2.08–2.33× as long as high. Submetapleural carina distinct, extended on anterior 0.3–0.65 of metapleuron. Propodeum shiny, with evenly sparse fine setiferous punctures on anterior half, in dorsal view 1.00–1.15× as long as medially wide. Propodeal spiracle 0.14–0.20 the length to anterior end of pleural carina. Hind leg with femur 3.20–3.70× as long as high, 0.97–1.0× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m 0.35–0.50× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.32–1.65× as long as Cu 1 b. Hind wing with vein cu-a + first abscissa of Cu 1 between M and cu-a slightly curved to angled, reclivous on ventral 0.3, vertical on dorsal 0.7; distal abscissa of Cu 1 absent. Metasoma (Fig. 37A, 37E, 37G, 37H). Tergite I 1.41–1.50× as long as posteriorly broad, shiny, with relatively dense setiferous punctures laterally and posteriorly, spiracle near its anterior 0.3–0.4; lateromedian longitudinal carinae present at anterior 0.2 of tergite, weak; lateral longitudinal carina complete, or interrupted medially, very weak. Sternite I extending posteriorly 0.55–0.7 the length of tergite. Tergite II 0.95–1.23× as long as posteriorly broad, with dense, wide and relatively shallow setiferous punctures, distance between punctures at most its diameter; remaining tergites same microsculpture. Ovipositor evenly up-curved at distal 0.3, 2.0–2.35× as long as hind tibia. Ovipositor sheath 1.6–1.8× as long as hind tibia, ventral margin basally without denticles, length of setae on average 1.96–2.04× the sheath basal width. Colouration (Fig. 37). Head mostly black with clypeus partially, facial and frontal orbits very narrowly insinuated, vertical orbits, spot behind mandibular basis, and mouth parts except apex of mandibles, white cream; clypeus partially brownish; antenna dark brown, pedicel, annellus and first and second flagellomeres, laterally whitish. Mesosoma mostly striking light orange; propleuron dorsally, anterior and posterior ventral part of pronotum, peripheral suture of mesoscutum, axillae, dorsal suture of mesopleuron, anterior and posterior margin of propodeum, ventral part of metapleuron, dark brown to black; dorsal lateral band on pronotum, tegula, postscutellum and lateral parts of propodeum, white cream. Metasoma mostly orange; sternite I, posterolateral spots on tergite I and posterior band on tergites II–V(VI), dark brown to black. Tergites I–V(VI) with white cream posterolateral spots just anteriorly to the dark band; ovipositor orange; ovipositor sheaths dark brown. Legs predominantly orange; front and mid leg with coxae, trochanters, trochantelli, anterior and distal part of femorae, anterior and posterior part of tibiae, and base of tarsi, whitish; hind leg with dorsal stripe on coxa, apex of trochanter, trochantellus, apex of femur, mid part of tibia and basal part of tarsi, white cream, mid and hind tibia and tarsi mostly dark brown, basal part of hind trochanter black. Wings hyaline, pterostigma blackish brown. Male: Body length 7.0– 7.7 mm. Fore wing length 4.9–5.0 mm. Head (Figs 38B, 38D–E, 38G). Gena, in dorsal view, 0.48–0.52× as long as eye. Posterior ocellus separated from eye 0.94–1.06× its maximum diameter. Distance between hind ocelli 1.02–1.24× the maximum diameter of posterior ocellus. Clypeus 1.68–1.80× as broad as medially long. Malar space 0.89–0.95× as long as basal mandibular width. Antenna with 24–25 flagellomeres, first flagellomere 4.66–5.02× as long as wide. Mesosoma (Figs 38A, 38B, 38G). Metapleuron 1.94–2.11× as long as high. Propodeum in dorsal view 1.07– 1.11× as long as medially wide. Propodeal spiracle 0.15–0.16 the length to anterior end of pleural carina. Hind leg with femur 3.32–3.89× as long as high, 0.83–0.92× as long as tibia. Fore wing with vein 2 rs-m 0.28–0.42× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.25–1.50× as long as Cu 1 b. Metasoma (Figs 38C, 38F, 38H). Setae on tergites longer and denser than in females. Tergite I 1.58–1.69× as long as posteriorly broad. Sternite I extending posteriorly 0.70–0.80 the length of tergite. Tergite II 1.38–1.53× as long as posteriorly broad. Colouration (Fig. 38). As in female but white cream facial orbits extending medially below antennal insertions, pronotum and propodeum extensively dark brown, hind coxa blackish laterally, and tergites brownish black, without white cream marks, only with slightly paler anterior lateral areas on tergites II and III. Other features as in female. Etymology. This species is named in honor to the Colombian indigenous tribe of Muiscas, who lived mainly in the territories that today correspond to the departments of Boyacá and Cundinamarca. Type material (13 ♀♀, 3 ♂♂). Holotype. Colombia: 1 ♀, Cundinamarca, Fontibón, Parque Centenario, 4 o 40'N, 74 o 08'W. 2549m, 31-VIII-2003, E. Ortiz (MJHN). Paratypes. Colombia: 1 ♀, Boyacá, Arcabuco, SFF Iguaque, Chaina, 05 o 25’N, 73 o 27’W, 2600 m, 29-I/ 14-II-2001, malaise trap, A. Roberto (IAvH); 4 ♀♀, same locality and trap type, 5–25-III-2001, P. Reina (IAvH); 2 ♀♀, same locality, collector and trap type, 26-III/ 12-IV-2001 (IAvH); 1 ♀, same locality, collector and trap type, 13-IV/ 5-V-2001 (IAvH); 1 ♀, 1 ♂, same locality, collector and trap type, 17-V/ 5-VI-2001 (IAvH); 1 ♀, same locality, collector and trap type, 6–25-VII-2001 (IAvH); 1 ♀, same locality, collector and trap type, 9–31-VIII-2001 (IAvH); 1 ♂, same locality, collector and trap type, 25-X/ 16- XI-2001 (IAvH); 1 ♂, same locality, collector and trap type, 1–16-XII-2001 (IAvH); 1 ♀, Cund [Cundinamarca], 2900m, Guasca, 10-III-1940, Chapin N. 694, USNM 2053570 (NMNH). Distribution. Colombia (Fig. 52B, Appendix I).