Clistopyga horcoana Bordera & Palacio & Herrera-Flórez 2025, sp. nov.

12. Clistopyga horcoana Bordera & Palacio, sp. nov. (Figs 18, 19, 52C, Appendix I).) Diagnosis. Clistopyga horcoana may be distinguished from all other species of the C. calixtoi species group by the following combination of characters: genal orbit interrupted by dark brown or black (Figs 18A, 18E, 19A, 19C); mesosoma extensively red orange (Figs 18C, 19C–D); pronotum mostly blackish brown and white cream (Figs 18C– D, 19C); propodeum laterally broadly white cream (Figs 18C, 19C); hind femur yellow with subbasal and subdistal lateral brown marks (Figs 18A, 19A); metasoma mostly striking light orange with posterior white cream bands on tergites II–V(VI in male) (Fig. 18F, 18H, 19G–H); tergite I, without wide white cream band (Figs 18F, 19G); epomia strong and short, vertical (Figs 18D, arrow, 19C); fore wing with vein 2 rs-m 0.2–0.4× as long as abscissa of M between 2 rs-m and 2 m-cu (Figs 18I, 19E); metapleuron 2.0–2.4× as long as high (Fig. 18G); propodeum in dorsal view 0.85–1.15× as long as medially wide (Fig. 19D). Additionally, female has submetapleural carina weak, present on anterior 0.5 of metapleuron (Fig. 18G); hind wing with distal abscissa of Cu 1 slightly insinuated (Fig. 18K); tergite I 1.25.1.35× as long as posteriorly broad (Fig. 18F); tergite II 0.9–1.0× as long as posteriorly broad, relatively deeply punctate (Fig. 18F); ovipositor 2.15–2.3× as long as hind tibia and moderately upcurved at distal 0.3 (Fig. 18J); ovipositor sheath 1.75–1.85× as long as hind tibia (Fig. 18A), ventral margin basally with small denticles. Male has metapleuron mostly reddish or orange (Fig.19C); gena, in frontal view, with a very deep semicircular concavity posterior to mandible (Fig. 19B); submetapleural carina strong, extended on anterior 0.6 of metapleuron; tergite II 1.45–1.46× as long as wide (Fig. 19G). Description. Female: Body length 5.6–8.6 mm. Fore wing length 4.1–6.1 mm. Head (Figs 18A–B, 18E). In dorsal view, strongly narrowed behind eyes. Gena shiny with very sparse setiferous punctures, in dorsal view slightly rounded, 0.26–0.28× as long as eye; in frontal view strongly constricted ventrally to eyes, slightly convex or straight. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 1.1–1.25× its maximum diameter. Distance between hind ocelli 1.1–1.25× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform behind mandible. Face with shallow, fine and moderately dense punctures, distance between them about its diameter. Clypeal suture slightly curved medially. Clypeus 2.1–2.3× as broad as medially long, weakly concave or straight, with very few setiferous punctures dorsally, flat and smooth ventrally, ventral margin straight centrally. Malar space 0.8–0.9× as long as basal mandibular width, with a smooth narrow subocular sulcus. Antenna with 23–26 flagellomeres, first flagellomere 6.5–7.0× as long as medially wide. Mesosoma (Figs 18A, 18C–E, 18G, 18I, 18K). Pronotum smooth and shiny with very few setiferous punctures on posterior corner. Epomia strong and short, vertical. Mesoscutum smooth and shiny, with moderately dense shallow and fine setiferous punctures, much denser on anterior part of median lobe; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus deep, reaching about 0.5 the length of mesoscutum. Mesopleuron smooth and shiny, with moderately dense small setiferous punctures on anterior and ventral part. Epicnemial carina weak, its dorsal end slightly curved towards the back, ending at level of centre of pronotum. Metapleuron smooth and shiny, with very shallow and isolated setiferous punctures, 2.0–2.4× as long as high. Submetapleural carina weak, present at anterior 0.5 of metapleuron. Propodeum shiny with evenly sparse fine setiferous punctures, in dorsal view 0.85–1.15× as long as medially wide. Propodeal spiracle 0.15–0.20× the length to anterior end of pleural carina. Hind leg with femur 3.25–3.5× as long as high, 1.0–1.05× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m 0.2–0.4× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.8–1.9× as long as Cu 1 b. Hind wing with vein cu-a 0.34–0.5 × as long as abscissa of Cu 1 between M and cu-a Vein cu-a reclivous, first abscissa of Cu 1 vertical; sometimes vein cu-a + abscissa of Cu 1 between M and cu-a not angled, slightly curved, reclival; distal abscissa of Cu 1 slightly insinuated. Metasoma (Figs 18A, 18F, 18H, 18J). Tergite I 1.25.1.35× as long as posteriorly broad, shiny, with moderately sparse shallow setiferous punctures laterally and posteriorly, spiracle near its anterior 0.4; lateromedian longitudinal carinae strong at anterior 0.3 of tergite; lateral longitudinal carinae strong, present at anterior 0.35 and posterior 0.2 of tergite. Sternite I extending posteriorly 0.47–0.5× the length of tergite. Tergite II 0.9–1.0× as long as posteriorly broad, with moderately deep setiferous punctures, distance between punctures more than its diameter. Ovipositor moderately up-curved at posterior 0.3, 2.15–2.3× as long as hind tibia. Ovipositor sheath 1.75–1.85× as long as hind tibia, ventral margin basally with small denticles, length of setae on average 2.0–2.5× the sheath basal width. Colouration (Fig. 18). Head mostly white cream, with apex of mandibles, frons and vertex medially, middle part of genal orbit, middle and dorsal posterior part of gena and occipital region, black; widely clypeal suture, clypeal fovea, subocular sulcus and longitudinal middle stripe on face, dark brown; antenna light brown, scapus, pedicel, annellus and first and second flagellomeres, laterally white cream. Mesosoma mostly light orange; propleuron dorsally, most part of pronotum, posterior dorsal part of mesopleuron, axillae, ventral part of metapleuron and posterior part of propodeum dark brown; propleuron ventrally, ventral anterior and dorsal lateral wide band on pronotum, two longitudinal stripes on mesoscutum, interrupted by notauli, a mid spot on anterior part of mesopleuron joining epicnemial carina, tegula, subalar prominence, scutellum laterally and posteriorly, postscutellum and lateral parts of propodeum, white cream. Metasoma mostly striking light orange; tergites II–V anterolaterally and posteriorly white cream banded, tergites I–III with lateral posterior black spots; ovipositor dark orange; ovipositor sheaths dark brown. Legs predominantly white cream; front leg with dorsal stripe on trochanter, femur, tibia and distal part of tarsi, brown; mid leg with base of trochanter, dorsal stripes on femur and tibia and distal part of tarsi brown; hind leg with anterior and posterior spots on coxa, base of trochanter, base, and lateral subdistal marks on femur, basal and distal part of tibia, and distal part of tarsi, brown. Wings hyaline, pterostigma blackish brown. Male: Body length about 6.4 mm. Fore wing length 3.7–3.8 mm. Head (Figs 19B–C). Gena about 0.33× as long as eye; in frontal view with a deep and conspicuous longitudinal concavity. Posterior ocellus separated from eye about 1.0 its maximum diameter. Distance between hind ocelli 0.95–1.1× maximum diameter of posterior ocellus. Clypeus 2.3–2.45× as broad as medially long. Malar space 0.8–1.0× as long as basal mandibular width. Antenna with 22 flagellomeres, first flagellomere 6.6–6.7× as long as medially wide. Mesosoma (Figs 19A, 19C–F). Metapleuron 2.0–2.2× as long as high. Submetapleural carina strong, present at anterior 0.6 of metapleuron. Propodeum in dorsal view 0.95–1.05× as long as medially wide. Propodeal spiracle about 0.16 the length to anterior end of pleural carina. Hind leg with femur 4.05–4.2× as long as high, 0.85–0.9× as long as tibia. Fore wing with vein cu-a slightly distal to Rs & M; vein 2 rs-m 0.25× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.35–1.45× as long as Cu 1 b. Hind wing with vein cu-a + abscissa of Cu 1 between M and cu-a not angled, reclival; distal abscissa of Cu 1 absent. Metasoma (Figs 19A, 19G–H). Tergite I 1.55–1.65× as long as posteriorly broad, shiny, with moderately dense shallow punctures, lateral longitudinal carina strong, present at posterior 0.2 of tergite. Tergite II 1.45–1.46× as long as posteriorly broad. Colouration (Fig. 19). As in female, but mesopleuron widely white cream ventrally. Other characters as in female. Etymology. The species name refers to the experimental reserve of Horco Molle in San Miguel de Tucuman (Argentina), where the holotype was collected. Type material (21 ♀♀, 2 ♂♂). Holotype. Argentina: 1 ♀, Horco Molle nr. Tucumán, 20–31-VIII-1966, Lionel Stange (EMUS). Paratypes. Argentina: 1 ♀, El Jardín, Salta, 2-X-1968, C. Porter (FSCA); 1 ♂, El Pintado, 650 m, 27–29-IX-1960, Cat., Luis Peña (EMUS); 1 ♀, Horco Molle nr Tucumán, 1–19-VIII-1966, Lionel Stange (EMUS); 3 ♀♀, same data, 20–31-VIII-1966 (EMUS); 2 ♀♀, same data,19.IX/ 7.X.1966 (EMUS); 1 ♀, same locality, 27.X.1967 (EMUS); 1 ♀, same locality, 15-XI-1967, C. Porter (FSCA); 1 ♀, same locality, 1–9-X-1968 (EMUS); 1 ♀, Huerta Grande [Cordoba], 2.IX.1959, Leg Hans Förster (SNSB-ZSM); 1 ♀, 11 Km W. Las Cejas, Tucumán, 27-V/ 14-VIII-1968, Lionel Stange (EMUS); 1 ♂, La Plata, 15-XII-1965, H & M Townes (EMUS); 5 ♀♀, La Rioja, Santa Cruz, 1700m, Humid Ravine, 1–15-I-2003. F. Fidalgo Malaise Trap (CEUA); 1 ♀, Provincia de Santiago del Estero, Bords du Rio Salado, Env. Dicano, E.R. Wagner 1909, decembre, janvier (MNHN); 1 ♀, Santiago del Estero, Termas de Rio Hondo, Dique frontal, 21-VIII-1976, C. Porter & L. Stange (FSCA); 1 ♀, Tucumán, Reserva Forestal, Rta. 9 Ca. El Cadillal, 20-VI-1973, C. Porter & E. Demarest (FSCA). Distribution. Argentina (Fig. 52C, Appendix I).