Clistopyga andaki Palacio & Bordera 2025, sp. nov.

2. Clistopyga andaki Palacio & Bordera, sp. nov. (Figs 4, 5, 52A, Appendix I) Diagnosis. Clistopyga andaki may be distinguished from other species of the C. calixtoi species group by the following combination of characters: genal orbits widely interrupted (Fig. 4D, 5C); mesopleuron mostly orange or reddish (Figs 4D, 5C); metapleuron and background of propodeum and metasomal tergites dark brown to black (Figs 4, 5); propodeum widely tinged yellow laterally (Figs 4D, 4G, 5C, 5H); face with fine and relatively sparse setiferous punctures (Fig. 4A, 5B); epomia as a small tubercle or short carina at the bottom of the submarginal depression of pronotum (Fig. 4F, 5E); submetapleural carina only present on anterior part (Fig. 4G, 5C). Additionally, female has metasoma with posterolateral well defined white cream marks on tergites, clearly separated from posterior rim (Fig. 4E); ventral margin of ovipositor sheath basally without denticles. Male with anterior part of mid lobe and lateral lobes of mesoscutum black (Fig. 5H); metasoma without posterolateral well defined white cream on tergites clearly separated from posterior rim (Figs 5F, 5I); gena without longitudinal concavity posterior to mandible base (Fig. 5B); tergite II 1.50× as long as posteriorly broad. Description. Female: Body length 5.4 mm. Fore wing length 4.0 mm. Head (Figs 4A, 4D). In dorsal view, moderately narrowed behind eyes. Gena shiny mostly smooth with uniformly sparse setiferous punctures, in dorsal view slightly rounded, 0.51× as long as eye, in frontal view moderately constricted below eyes, slightly convex. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 1.23× its maximum diameter. Distance between hind ocelli 1.27× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform behind mandible. Face with fine and relatively sparse setiferous punctures, distance between punctures more than twice the diameter of punctures. Clypeal suture strongly curved. Clypeus 1.82× as broad as medially long, distinctly convex in dorsal half, flat in ventral half, with apical margin slightly concave. Malar space 0.97× as long as basal mandibular width, with a weak, slightly granulate subocular sulcus. Antenna with 24 flagellomeres, first flagellomere 5.56× as long as wide. Mesosoma (Figs 4B–D, 4F–H). Pronotum shiny, smooth, with fine and scattered setiferous punctures in dorsal posterior part. Epomia as short carina at the bottom of the submarginal depression of pronotum. Mesoscutum entirely smooth and shiny, with fine and dense setiferous punctures; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus moderately deep, reaching about 0.5 the length of mesoscutum. Mesopleuron smooth and shiny, with evenly sparse setiferous punctures on anterior and ventral part. Epicnemial carina strong, its dorsal end straight, ending at level of centre of pronotum. Metapleuron shiny, glabrous, 2.46–2.47× as long as high. Submetapleural carina weak, extended on anterior 0.4–0.5 of metapleuron. Propodeum smooth and shiny, with very sparse fine setiferous punctures on anterior dorsal part and laterally, in dorsal view 1.40× as long as medially wide. Propodeal spiracle about 0.16 the length to anterior end of pleural carina. Hind leg with femur about 3.0× as long as high, 0.94× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m about 0.43× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.20× as long as Cu 1 b. Hind wing with vein cu-a + first abscissa of Cu 1 between M and cu-a not angled, slightly curved, reclivous; distal abscissa of Cu 1 absent. Metasoma (Figs 4E, 4G, 4I–J). Tergite I 1.85× as long as posteriorly broad, shiny, with sparse setiferous punctures laterally and posteriorly; spiracle near its anterior 0.4; lateromedian longitudinal carinae present at anterior 0.2 of tergite, very weak; lateral longitudinal carinae present at posterior 0.3 of tergite. Sternite I extending back about 0.7 of length of tergite. Tergite II 1.50× as long as posteriorly broad, with dense, shallow, wide setiferous punctures, distance between punctures less than its diameter; punctation of posterior tergites tending to be deeper and less wide. Ovipositor relatively stout, evenly up-curved at distal 0.35, 2.1× as long as hind tibia. Ovipositor sheath about 1.7× as long as hind tibia, ventral margin basally without denticles, length of setae on average about 2.15× the sheath basal width. Colouration (Fig. 4). Head black with mouthparts, except apex of mandibles, clypeus, facial orbits weakly, extending below antennal sockets, frontal and vertical orbits, and ventral part of gena, white cream; antenna brownish, ventrally pallid towards base, pedicel and apex of scape ventrally white. Mesosoma with lateral lobes of mesoscutum, scutellum, mesopleuron, mesosternum and most of mesepimeron reddish; ventral part of propleuron, anterior and dorsal margin of pronotum, posterior part of scutellum, postscutellum, tegula, dorsal part of mesepimeron and sides of propodeum white cream; median lobe of mesoscutum, most of propleuron, most of pronotum, anterior, dorsal, and posterior part of propodeum, and metapleuron brownish black. Metasoma black to blackish brown; tergites II–V posteriorly white marked at sides, tergites I–IV with posterolateral margins black marked; ovipositor dark brown; ovipositor sheath black. Anterior two pair of legs predominantly white, fore legs with femora, tibia and tarsi slightly infuscate, middle legs with trochantellus proximally black, femora and tibia dorsally striped with black and tarsi infuscate; hind leg white with coxa laterally, trochanter proximally, femur proximally and subdistally, tibia proximally and distally, and first tarsomere distally black; remain tarsal segments blackish. Wings hyaline, pterostigma blackish brown. Male: Body length 6.1–8.1 mm. Fore wing length 4.8–5.3 mm. Head (Figs 5B–D). Gena, in dorsal view, somewhat rounded, 0.38× as long as eye, without an impressed longitudinal groove behind mandibular base. Posterior ocellus separated from eye 0.98× its maximum diameter. Distance between hind ocelli 0.86× the maximum diameter of posterior ocellus. Clypeus 1.51× as broad as medially long. Malar space 0.69× as long as basal mandibular width. Antenna with 26 flagellomeres, first flagellomere 5.6–5.7× as long as wide. Mesosoma (Figs 5C, 5E, 5G–H). Metapleuron 2.44× as long as high. Propodeum in dorsal view 1.35–1.38× as long as medially wide. Propodeal spiracle about 0.18 the length to anterior end of pleural carina. Hind leg with femur 4.5–4.6× as long as high, about 0.80× as long as tibia. Fore wing with vein 2 rs-m 0.33–0.36× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.12× as long as Cu 1 b. Metasoma (Figs 5F, 5I). Setae of tergites longer and denser than in females. Tergite I 2.03–2.06× as long as posteriorly broad. Sternite I extending back 0.6 the length of tergite. Tergite II 1.65–1.70× as long as posteriorly broad. Colouration (Fig. 5). As in female but with mesosoma black in the areas that are blackish brown in the female, and with metasomal tergites entirely black. Other features as in female. Remarks. The three males assigned to this species are larger than the female and were collected in another locality, but are very similar in structure and colour pattern, so they are tentatively assigned to C. andaki. Etymology. This species is named in honor to the Colombian indigenous tribe of Andakis (or Andaquíes), who live in southern Dept. of Huila, where the holotype was collected. Type material (1 ♀, 3 ♂♂). Holotype. Colombia: 1 ♀, Huila, PNN Cueva de los Guácharos, Alto el Mirador, 1 º 38'N, 76 º 6'W, 1980 m, 21-IV/ 5-V-2002, malaise trap, J. Fonseca (IAvH). Paratypes. Colombia: 1 ♂, Valle, PNN Farallones de Cali, Los Andes, vda Quebradahonda, 3 º 34'N, 76 º 40'W, 1730 m, 15-XII-1998, N. Beltrán (IAvH); 1 ♂, same locality and collector, 30-XII-1998 (IAvH); 1 ♂, same locality, 23-XII-1998, W. Alfonso (IAvH). Distribution. Colombia (Fig. 52A, Appendix I).