Euura vaga

303 Euura vaga (Fabricius, 1781) Figs 146, 542 Tenthredo vaga Fabricius, 1781: 415. Syntypes (3 specimens in NHMD according to Zimsen 1964), sex not stated, not examined. Type locality: England. Tenthredo flaviventris Gmelin, 1790: 2667. Syntypes, sex not stated, not located. Type locality: Europe. Synonymy by Konow (1904c). Primary homonym of Tenthredo flaviventris Retzius, 1783 [Neurotoma saltuum (Linnaeus, 1758)]. Nematus leucogaster Hartig, 1840: 23. Lectotype designated below. Synonymy by Konow (1904c). Nematus hypoleucus Förster, 1854a: 304–305. Lectotype designated below. Synonymy by Konow (1904c). Nematus micraulius Förster, 1854b: 428–429. Lectotype designated below. Synonymy by Konow (1904c). Nematus punctipleuris Thomson, 1863: 628. Lectotype designated below. Synonymy by Konow (1904c). Nematus punctulatus Thomson, 1863: 626–627. Lectotype designated below. Synonymy by Konow (1904c). Nematus inconspicuus W.F. Kirby, 1882: 141. Holotype ♀ (BMNH?), not examined. Type locality: New York. Synonymy by Benson (1962). Pachynematus corticosus MacGillivray, 1901: 584–585. Holotype ♀ (INHS), not examined. Type locality: USA, New York, Saranac Inn. Holcocneme ulbrichti Enslin, 1910: 316–317. Holotype ♀, not found in ZSM. Type locality: Germany, Nordrhein-Westfalen, Krefeld. Synonymy by Enslin (1915). Pachynematus roscidus MacGillivray, 1921b: 31. Holotype ♀ (INHS), not examined. Type locality: USA, Maine, Orono. Pachynematus rugosulus Lindqvist, 1959b: 71. Syn. nov. Diagnosis The female is most similar to E. obducta from which it differs in usually having a completely pale metafemur (extensively black in E. obducta), more triangular valvula 3 in dorsal view, and apically curved basal part of lamnium (more or less straight in E. obducta). Males, which are extremely rare in Europe, are best recognised by their penis valves. Abdomen sometimes also dorsally nearly completely pale (terga 3–10) (ZMUO.062019). Pterostigma varies in European specimens from yellowish to dark brown, in North America often (usually?) black. Type material examined Lectotype Nematus leucogaster, here designated GERMANY • ♀; northern Germany [according to Hartig 1840: 22]; ZSM, GBIF-GISHym3334. Lectotype Nematus hypoleucus, here designated GERMANY – Nordrhein-Westfalen • ♀; Aachen area; 50.77° N, 6.09° E; ZSM, GBIF-GISHym3297. Lectotype Nematus micraulius, here designated GERMANY – Nordrhein-Westfalen • ♂; Aachen area; 50.77° N, 6.09° E; ZSM, GBIF-GISHym3352. Notes Penis valves lost, but identified by Conde as Pristiphora vaga. Lectotype Nematus punctipleuris, here designated SWEDEN • ♀; Norlandia [Lapland, probably Sweden]; MZLU, MZLU2017272. Paralectotype Nematus punctipleuris SWEDEN • ♀; Norlandia [Lapland, probably Sweden]; MZLU, MZLU2017273. Lectotype Nematus punctulatus, here designated SWEDEN • ♀; Torne Lappmark; MZLU, MZLU2017265 Paralectotypes Nematus punctulatus SWEDEN • 2 ♀♀; Torne Lappmark; MZLU, MZLU 2017266, MZLU 2017269 • 2 ♀♀; [labels illegible]; MZLU, MZLU 2017267, MZLU 2017268. Holotype Pachynematus rugosulus NORWAY – Finnmark • ♀; Jotkajavrek; 69.77° N, 23.85° E; Jul. 1924; Expedition A. Strand leg.; ZMUN, NHMO:1000201217. Notes Basal quarter to third of metafemur brown, mesepisternum matt. Best morphological match: ZMUO.041887. Host plants Carex spp. More than one generation per year in suitable conditions (Lorenz & Kraus 1957; Kangas 1985). Genetics COI Based on 29 specimens, maximum within-species distance is 3.5% and the nearest neighbour, diverging by a minimum of 6.26%, is Euura miliaris. Including 8 specimens from North America does not change the results. Nuclear Based on 8 specimens, maximum within-species distance is 0.61% (1.02% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 2.93%, is Euura fuscomaculata. When a specimen from North America (DEI-GISHym15293) is included, maximum within-species distance is 0.8% (1.02% based on haplotypes of individual females). Distribution and material examined Holarctic. Specimens studied are from Canada, Estonia, Finland, France, Germany, Iceland, Norway, Russia, Slovakia, Sweden, United Kingdom, and the USA.